PBDB Collection

Tsagan Khushu [HMNS-GIN] (Cretaceous of Mongolia)

Also known as Tsagan Khusku, Tsagan Usu, Tsagan Hushu

Where: Omnogov, Mongolia (43.5° N, 100.4° E: paleocoordinates 41.1° N, 89.4° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: Nemegt Formation, Maastrichtian (72.1 - 66.0 Ma)

• formation-level stratigraphic resolution

Environment/lithology: fluvial; fine-grained, carbonaceous sandstone and mudstone

• "The bituminous sandstone with fine horizontal lamination was found in the part of alternation of mudstone and sandstone"; "The fossiliferous beds of this area are alternation of sandstone and mudstone layers with fluvial origin. Those beds are horizontally situated. Dinosaur bones and associated skeletons are included in those layers. There is a finely stratified mudstone lens with high contents of carbon material that yield pollen and spore."

Size classes: macrofossils, mesofossils, microfossils

Collected by HMNS-GIN in 1993, 1997, 2006

Collection methods: bulk, surface (float), surface (in situ),

• HMNS-Hayashibara Museum of Natural Sciences

Primary reference: M. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural Sciences Research Bulletin 1:17-29 [R. Benson/R. Benson/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 127681: authorized by Roger Benson, entered by Roger Benson on 22.05.2012, edited by Matthew Carrano

Creative Commons license: CC BY (attribution)

Taxonomic list

• JMJPE researched here in 1993 and 1997, and collected one partial skull of a pachycephalosaur, fragments of skull elements of Tarbosaurus, petrified wood, and several forms of dinosaur eggshells. "Pollen and spore fossils" found as well.

Show authors, comments, and common names

Angiospermae
-	Angiospermae indet.⁷ Angiospermae indet.⁷ pollen

Tracheophyta -	Tracheophyta indet.⁵ Tracheophyta indet.⁵ 97076 TK Johnson; "petrified wood"
Reptilia
Dinosauria -	Dinosauria indet. Dinosauria indet. Owen 1842 dinosaur digits
Theropoda -	Oviraptorosauria indet.⁶ Oviraptorosauria indet.⁶ Barsbold 1976 maniraptoran
Theropoda - Tyrannosauridae	Tarbosaurus sp. Tarbosaurus sp. Maleev 1955 tyrannosaurid skull elements of Tarbosaurus
Theropoda - Dendroolithidae	Dendroolithidae indet.⁶ Dendroolithidae indet.⁶ Zhao and Li 1988 coelurosaur
Theropoda - Ornithomimidae	Gallimimus sp. Gallimimus sp. Osmólska et al. 1972 ornithomimid femur
Theropoda - Elongatoolithidae	Elongatoolithidae indet.⁶ Elongatoolithidae indet.⁶ Zhao 1975 theropod
Ornithischia -	Ankylosauria indet.⁵, Prenocephale sp.¹ Ankylosauria indet.⁵ Osborn 1923 ankylosaur Prenocephale sp.¹ Maryańska and Osmólska 1974 ornithischian 930918 TK; partial skull = MPC-D 100/1206
Ornithischia - Spheroolithidae	Spheroolithidae indet.⁶ Spheroolithidae indet.⁶ Zhao 1979 ornithopod
Ornithischia - Hadrosauridae	Saurolophus angustirostris⁴ Saurolophus angustirostris⁴ Rozhdestvensky 1952 hadrosaurine 060903 TK TSGT
Saurischia -	Sauropoda indet.⁵ Sauropoda indet.⁵ Marsh 1878 sauropod A right ilium, femur, tibia and fibula of a sauropod
Crocodylia -	Crocodylia indet. Crocodylia indet. Owen 1842 crocodilian scute
Testudinata - Sichuanchelyidae	Mongolochelys efremovi³ Mongolochelys efremovi³ Khosatzky 1997 turtle
Testudines - Lindholmemydidae	Mongolemys sp.² Mongolemys sp.² Khosatzky and Mlynarski 1971 turtle 060905 TK TSI
Testudines - Trionychidae	Trionychidae indet.² Trionychidae indet.² Gray 1825 softshell turtle

Mollusca -	Mollusca indet.⁵ Mollusca indet.⁵ mollusk

¹Evans et al. 2018; ²Suzuki and Narmandakh 2004; ³Suzuki and Tsogtbaatar 2010; ⁴Tsogtbaatar and Chinzorig 2007; ⁵Watabe and Suzuki 2000; ⁶Watabe et al. 2010; ⁷Watabe et al. 2010