Shanwang, Linqu County, collection 79/80 (Beijing Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...

Primary reference: Y. C. Hong. 1983. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128550: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012, edited by Erin Leckey

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Fulgoridae
Aphaena lithoecia n. sp.5, "Oxycephala xiejiaheensis" = Limois shanwangensis5, "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis
Aphaena lithoecia n. sp.5 Zhang 1989 fulgorid planthopper
79022
"Oxycephala xiejiaheensis" = Limois shanwangensis5 Hong 1979 fulgorid planthopper
79026, 79079, 79080
"Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis Hong 1979 fulgorid planthopper
79080
 Hemiptera - Aphididae
"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia5
"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia5 Zhang 1989 aphid
79056
 Hemiptera - Cicadidae
"Oncotympana lapidescens n. sp." = Hyalessa lapidescens5
"Oncotympana lapidescens n. sp." = Hyalessa lapidescens5 Zhang 1989 cicada
79023
 Hemiptera -
Heteroptera indet.5 Latreille 1810 true bug
79086
 Hemiptera - Coreidae
"Coriopsis miocenica n. sp." = Coreiopsis miocenica3, Coreiopsis punctata n. gen. n. sp., "Coriopsis parva n. sp." = Coreiopsis parva3, Shanwangicoris longa n. gen. n. sp.3, Gonocerus shandongensis n. sp.5, Cloresmus shanwangi n. sp.5
"Coriopsis miocenica n. sp." = Coreiopsis miocenica3 Hong and Wang 1987 leaf-footed bug
80104
Coreiopsis punctata n. gen. n. sp. Hong 1983 leaf-footed bug
79084
"Coriopsis parva n. sp." = Coreiopsis parva3 Hong and Wang 1987 leaf-footed bug
80115
Shanwangicoris longa n. gen. n. sp.3 Hong and Wang 1987 leaf-footed bug
80129
Gonocerus shandongensis n. sp.5 Zhang 1989 leaf-footed bug
79025
Cloresmus shanwangi n. sp.5 Zhang 1989 leaf-footed bug
79005
 Hemiptera - Urostylididae
Urochela melaina n. sp.5, "Coreiopsis punctatus n. sp." = Urochela maculosa5
Urochela melaina n. sp.5 Zhang 1989 true bug
79024
"Coreiopsis punctatus n. sp." = Urochela maculosa5 Zhang 1989 true bug
79084
 Hemiptera - Nepidae
Coroides xiejiaheense n. gen. n. sp.
Coroides xiejiaheense n. gen. n. sp. Hong 1983 true bug
79113
 Hemiptera - Belostomatidae
Manocerus stagnans n. gen. n. sp.5
Manocerus stagnans n. gen. n. sp.5 Zhang 1989 giant water bug
79016
 Hemiptera - Notonectidae
Tarsonecta mecopoda n. gen. n. sp.5
Tarsonecta mecopoda n. gen. n. sp.5 Zhang 1989 true bug
79013
 Dermaptera - Forficulidae
Apanechura asecta n. sp.5 Zhang 1989 common earwig
79029
 Hymenoptera - Vespidae
"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis
"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis Hong 1983 potter wasp
79044
 Hymenoptera - Tiphiidae
"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis
"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis Hong 1983 wasp
79041
 Hymenoptera - Formicidae
? Lasius truncatus n. sp.5, Lasius vetulus5, Lasius validus n. sp.5, Formica linquensis n. sp.5, "Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis, Camponotus curviansatus n. sp.5, Camponotus microthoracus n. sp.5, "Shanwangella palaeoptera n. gen. n. sp." = Camponotus palaeopterus5, Camponotus longus n. sp.5, Camponotus mitis5, "Camponotus shanwangense" = Camponotus shanwangensis, "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5, Liometopum potamophilum n. sp.5, ? Tapinoma baculum n. sp.5, "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5, Paraphaenogaster shanwangensis n. sp.
? Lasius truncatus n. sp.5 Zhang 1989 ant
79055
Lasius vetulus5 Dlussky 1981 ant
79030
Lasius validus n. sp.5 Zhang 1989 ant
79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4)
Formica linquensis n. sp.5 Zhang 1989 ant
79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4)
"Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis Hong 1983 ant
79069
Camponotus curviansatus n. sp.5 Zhang 1989 ant
79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1)
Camponotus microthoracus n. sp.5 Zhang 1989 ant
79046, 79127, 79128
"Shanwangella palaeoptera n. gen. n. sp." = Camponotus palaeopterus5 Zhang 1989 ant
79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2)
Camponotus longus n. sp.5 Zhang 1989 ant
79052
Camponotus mitis5 Smith 1892 ant
79067 (Apis miocenica in Hing, 1985 - pl 22, fig 3)
"Camponotus shanwangense" = Camponotus shanwangensis Hong 1983 ant
79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985
"Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5 Hong 1985 ant
79087; 80196 in Hong & Wang 1985
Liometopum potamophilum n. sp.5 Zhang 1989 ant
79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6)
? Tapinoma baculum n. sp.5 Zhang 1989 ant
79051
"Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5 Zhang 1989 ant
79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3)
Paraphaenogaster shanwangensis n. sp. Hong 1983 ant
79047
 Hymenoptera - Apidae
Apis miocenica n. sp. Hong 1983 Chinese Miocene honey bee
79040, 79050; 80133 in Hong & Wang 1985
 Hymenoptera - Pemphredonidae
"Sinostigma spinalata n. sp." = Pemphredon spinalatum
"Sinostigma spinalata n. sp." = Pemphredon spinalatum Hong 1983 aphid wasp
79091
 Hymenoptera - Thynnidae
Elis pyrula n. sp.5
Elis pyrula n. sp.5 Zhang 1989 wasp
79089
 Hymenoptera - Ichneumonidae
Luia lacaris n. gen. n. sp.5
Luia lacaris n. gen. n. sp.5 Hong 1985 ichneumon wasp
79037
 Hymenoptera - Xyelidae
"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis
"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis Hong 1983 sawfly
79048
 Hymenoptera - Cimbicidae
"Procimbex shandongense n. sp." = Abia shandongensis1, "Procimbex shanwangensis n. sp." = Abia shanwangensis, Abia paurocephala n. sp.5
"Procimbex shandongense n. sp." = Abia shandongensis1 Hong and Wang 1985 cimbicid sawfly
80193
"Procimbex shanwangensis n. sp." = Abia shanwangensis Hong 1983 cimbicid sawfly
79038
Abia paurocephala n. sp.5 Zhang 1989 cimbicid sawfly
79085
 Coleoptera -
Polyphaga indet.5 Emery 1886 beetle
79063
 Coleoptera - Buprestidae
Procarabus punctatus n. gen. n. sp., Agrilus corrugatus n. sp.5
Procarabus punctatus n. gen. n. sp. Hong 1983 jewel beetle
79118/9
Agrilus corrugatus n. sp.5 Zhang 1989 jewel beetle
79062
 Coleoptera - Dascillidae
Dascillus elongatus5 Zhang 1989 soft bodied plant beetle
79081
Dascillus shandongianus5 Zhang 1989 soft bodied plant beetle
79009, 79017, 79077, 79088
 Coleoptera - Staphylinidae
Lathrobium homorum n. sp.5 Zhang 1989 rove beetle
79021
 Coleoptera - Meloidae
? Epicauta rufithoraca n. sp.5 Zhang 1989 blister beetle
79072
 Coleoptera - Tenebrionidae
Sinocistela siphla n. gen. n. sp.5
Sinocistela siphla n. gen. n. sp.5 Zhang 1989 comb-clawed beetle
79028
 Coleoptera - Endomychidae
"Engonius alviolatus" = Gerstaeckerus alviolatus5
"Engonius alviolatus" = Gerstaeckerus alviolatus5 Hong 1983 handsome fungus beetle
79041
 Coleoptera - Cerambycidae
Cerambycidae "indet. 1"5 Latreille 1802 longhorned beetle
79020
Mesocacia pulla n. sp.5 Zhang 1989 longhorned beetle
79019
Sinocalosoma longicornuta n. sp.2 Hong and Wang 1986 longhorned beetle
80175
Mesosa soteria n. sp.5 Zhang 1989 longhorned beetle
79008
Alifasciola daedalea n. sp.5 Zhang 1989 longhorned beetle
79074
 Coleoptera - Curculionidae
"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2, "Sinocurculia granulata n. sp." = Atactogaster granulata, Hylobius medianus n. sp.5, Hylobius aff. niitakensis5, Hylobius plenus5
"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2 Hong 1985 snout beetle
80128, 78001, 80107, 80134, 80173, 80174
"Sinocurculia granulata n. sp." = Atactogaster granulata Hong 1983 snout beetle
79109
Hylobius medianus n. sp.5 Zhang 1989 snout beetle
79117
Hylobius aff. niitakensis5 Kono 1933 snout beetle
79106, 79116 (ssp. fukienensis)
Hylobius plenus5 Zhang 1989 snout beetle
79010, 79012, 79073
 Coleoptera - Brentidae
Longifistulia parva n. gen. n. sp.3
Longifistulia parva n. gen. n. sp.3 Hong and Wang 1987 weevil
80116
 Coleoptera -
Scarabaeoidea "indet. 1"5 Latreille 1802 scarab beetle
79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2)
Aliscarabaeus granulatus n. sp. Hong 1983 scarab beetle
79111
 Coleoptera - Scarabaeidae
Holotrichia spatha n. sp.5, Anomala sp.5, "Magniscarabaeus furvus" = Anomala furva2, "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5, Anomala synemosyna n. sp.5, Anomala lochmocola n. sp.5
Holotrichia spatha n. sp.5 Zhang 1989 scarab beetle
79102
Anomala sp.5 Samouelle 1819 scarab beetle
79096
"Magniscarabaeus furvus" = Anomala furva2 Hong 1983 scarab beetle
80116, 80143
"Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5 Krell 2000 scarab beetle
79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987
Anomala synemosyna n. sp.5 Zhang 1989 scarab beetle
79006, 79007
Anomala lochmocola n. sp.5 Zhang 1989 scarab beetle
79112
 Coleoptera - Geotrupidae
Geotrupidae indet.5, "Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense
Geotrupidae indet.5 Latreille 1802 earth-boring dung beetle
79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3)
"Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Hong 1983 earth-boring dung beetle
79114/5
 Coleoptera - Carabidae
? Carabus ovalus n. sp.2, Calosoma cf. maderae5, "Shanwangicarabus furvus n. sp." = Chlaenius furvus2, "Shanwangicarabus paucinus n. sp." = Chlaenius furvus2, Scarites mancus n. sp.5
? Carabus ovalus n. sp.2 Hong and Wang 1986 ground beetle
80192
Calosoma cf. maderae5 Fabricius 1775 ground beetle
79001, 79002, 79108
"Shanwangicarabus furvus n. sp." = Chlaenius furvus2 Hong 1983 ground beetle
80125, 80106, 80126, 480183
"Shanwangicarabus paucinus n. sp." = Chlaenius furvus2 Hong 1983 ground beetle
80131
Scarites mancus n. sp.5 Zhang 1989 ground beetle
79106 (Aliscarabaeus granulatus in Hong, 1985)
 Coleoptera -
"Procarabus longipes n. sp." = Cistelites longipes5
"Procarabus longipes n. sp." = Cistelites longipes5 Hong 1985 beetle
79110
 Coleoptera - Cupedidae
Cupes longus n. sp.4
Cupes longus n. sp.4 Hong and Wang 1987 reticulated beetle
BNHM 80109
 Diptera - Bibionidae
Plecia solaris n. sp.5, Plecia ludongensis n. sp.1, Clothonopsis miocenica n. gen. n. sp.4, "Bibio miocenica n. sp." = Bibio miocenicus1, Bibio ruptineurus n. sp.5, Bibio cratopodus5, Bibio expansus5, Bibio deformatus n. sp.5, Bibio brunneus n. sp.5, Bibio shanwangensis1, "Fungivora bullata n. sp." = Bibio bullatus5, Bibio ventricosus n. sp.5
Plecia solaris n. sp.5 Zhang 1989 love bug
79125/6 (Bibio shanwangensis in Hong, 1985 - fig 26)
Plecia ludongensis n. sp.1 Hong and Wang 1985 love bug
80170
Clothonopsis miocenica n. gen. n. sp.4 Hong and Wang 1987 love bug
BNHM 80195
"Bibio miocenica n. sp." = Bibio miocenicus1 Hong and Wang 1985 march fly
80117
Bibio ruptineurus n. sp.5 Zhang 1989 march fly
79123 (Bibio sp. in Hong, 1985)
Bibio cratopodus5 Zhang 1989 march fly
79092
Bibio expansus5 Zhang 1989 march fly
79053 (Bibio shanwangensis in Hong, 1985 - pl 7, fig 4)
Bibio deformatus n. sp.5 Zhang 1989 march fly
79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1)
Bibio brunneus n. sp.5 Zhang 1989 march fly
79120 (Bibio sp. in Hong, 1985)
Bibio shanwangensis1 Hong 1983 march fly
80123
"Fungivora bullata n. sp." = Bibio bullatus5 Hong 1985 march fly
79003
Bibio ventricosus n. sp.5 Zhang 1989 march fly
79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3)
 Diptera - Sciaridae
Sciara brevipetiola n. sp.5 Zhang 1989 dark-winged fungus gnat
79004
 Diptera - Tipulidae
Tipula insolita n. sp.5 Zhang 1989 crane fly
79027
 Ephemeroptera - Leptophlebiidae
Paraleptophlebia sp.5 Lestage 1917 mayfly
79129 (Baetis sp. in Hong, 1985)
 Ephemeroptera - Heptageniidae
? Heptagenia sp.5 Walsh 1863 stream mayfly
79095
 Odonata - Gomphaeschnidae
Jagoria sinica n. sp.5 Zhang 1989 true dragonfly
79064 (Nasiaeschna orientalis in Hong, 1985)
Arachnida
 Araneae - Sparassidae
"Retina robusta" = Heteropoda robusta6
"Retina robusta" = Heteropoda robusta6 Hong 1985 spider
79104
 Araneae - Araneidae
Araneus piceus6, Araneus ruidipedalis n. sp.6, "Melanites orbiculatus" = Tethneus orbiculatus6
Araneus piceus6 Lin et al. 1989 orb-weaving spider
79099 (part of M. orbiculatus in Hong, 1985)
Araneus ruidipedalis n. sp.6 Zhang et al. 1994 orb-weaving spider
79098
"Melanites orbiculatus" = Tethneus orbiculatus6 Hong 1985 orb-weaving spider
79100
 Araneae - Tetragnathidae
"Eocryphoeca parva" = Tetragnatha parva6
"Eocryphoeca parva" = Tetragnatha parva6 Hong 1985 stretch spider
79101