Shanwang, Linqu County, collection 3 (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Provincial Museum (LPM). This collection contains specimens labeled with six-digit numbers starting with 8......

Primary reference: J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128551: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Arachnida
 Araneae - Thomisidae
"Ectona transipeda" = Ecotona transipeda2, Miothomisus sylvaticus n. gen. n. sp.2
"Ectona transipeda" = Ecotona transipeda2 Lin et al. 1989 crab spider
820175
Miothomisus sylvaticus n. gen. n. sp.2 Zhang et al. 1994 crab spider
820188
Insecta
 Blattaria - Phyllodromiidae
Phantocephalus meridionalis n. gen. n. sp. Zhang 1989 cockroach
830088
 Dermaptera - Forficulidae
Anechura cf. japonica de Bormans 1880 common earwig
840096
Allodahlia shanwangensis n. sp. Zhou 1986 common earwig
820156, 830113, 840004, 840074, 840117
Apanechura hydrophila n. gen. n. sp. Zhang 1989 common earwig
820130, 820068
Apanechura ooidea n. sp. Zhang 1989 common earwig
820174
 Blattodea - Blattidae
Periplaneta hylecoeta n. sp. Zhang 1989 cockroach
840134
 Orthoptera - Tettigoniidae
Tettigoniidae indet. Krauss 1902 katydid
820026
Brachypoda stagna n. gen. n. sp. Zhang 1989 katydid
820025
 Orthoptera - Acrididae
Aestilocusta graminea n. gen. n. sp.
Aestilocusta graminea n. gen. n. sp. Zhang 1989 band-winged grasshopper
820090
 Coleoptera - Scarabaeidae
Petraeianus ruderatus n. gen. n. sp., Anomala eversa n. sp., Adoretus recticlypeus n. sp., Mioserica margelis n. gen. n. sp., Macronota shandongiana n. sp., Genonota scola n. sp., Genonota mochthera n. gen. n. sp.
Petraeianus ruderatus n. gen. n. sp. Zhang 1989 scarab beetle
840089
Anomala eversa n. sp. Zhang 1989 scarab beetle
830124
Adoretus recticlypeus n. sp. Zhang 1989 scarab beetle
820192
Mioserica margelis n. gen. n. sp. Zhang 1989 scarab beetle
830089
Macronota shandongiana n. sp. Zhang 1989 flower chafer
840121
Genonota scola n. sp. Zhang 1989 flower chafer
830125
Genonota mochthera n. gen. n. sp. Zhang 1989 flower chafer
820129
 Coleoptera - Staphylinidae
Sinolispinodes torosus n. gen. n. sp. Zhang 1989 rove beetle
840059
Algon spatiosus n. sp. Zhang 1989 rove beetle
820145
Lathrobium sobrinum n. sp. Zhang 1989 rove beetle
830144
 Coleoptera - Dascillidae
Dascillus relictus n. sp. Zhang 1989 soft bodied plant beetle
840023
Dascillus flatus Zhang 1989 soft bodied plant beetle
820201, 840062
Dascillus shandongianus n. sp. Zhang 1989 soft bodied plant beetle
820028, 820049, 820167, 820219, 820231, 830116, 830084, 840064, 840890, 840892
Dascillus elongatus n. sp. Zhang 1989 soft bodied plant beetle
820243, 840069
Lyprodascillus sypharus n. gen. n. sp. Zhang 1989 soft bodied plant beetle
840068
 Coleoptera - Elateridae
Elater canabinus n. sp. Zhang 1989 click beetle
840105
 Coleoptera - Cantharidae
"Cantharis guttula n. sp." = Lycocerus guttula, "Curticantharis capacis n. gen. n. sp." = Themus (Themus) capacis
"Cantharis guttula n. sp." = Lycocerus guttula Zhang 1989 soldier beetle
830063
"Curticantharis capacis n. gen. n. sp." = Themus (Themus) capacis Zhang 1989 soldier beetle
820289
 Coleoptera - Buprestidae
Poecilonota nitela n. sp. Zhang 1989 jewel beetle
820253
Eolampra brachyelytra n. gen. n. sp. Zhang 1989 jewel beetle
840114
 Coleoptera - Vesperidae
Philus brachyelytrus n. sp. Zhang 1989 beetle
840094
 Coleoptera - Cerambycidae
? Cerambycidae "indet. 2" Latreille 1802 longhorned beetle
820263
Saxosa fusina n. gen. n. sp. Zhang 1989 longhorned beetle
840045
Eutetrapha striolata n. sp. Zhang 1989 longhorned beetle
820279
Mesosa laxa n. sp. Zhang 1989 longhorned beetle
820001
 Coleoptera - Chrysomelidae
? Smaragdina incerta n. sp. Zhang 1989 leaf beetle
830019
 Coleoptera - Curculionidae
Curculionidae indet. Latreille 1802 snout beetle
840079
Hylobius plenus n. sp. Zhang 1989 snout beetle
830112, 830139
 Coleoptera - Attelabidae
? Oedeuops instabilis n. sp. Zhang 1989 leaf rolling weevil
830078
 Coleoptera - Anthribidae
Tropideres fossilis n. sp. Zhang 1989 fungus weevil
820008
 Coleoptera - Coccinellidae
Coccinellidae indet. Latreille 1807 ladybird beetle
830039
 Coleoptera - Endomychidae
"Engonius alviolatus" = Gerstaeckerus alviolatus
"Engonius alviolatus" = Gerstaeckerus alviolatus Hong 1984 handsome fungus beetle
840067
 Coleoptera - Cleridae
Thanasimus taenianus n. sp. Zhang 1989 checkered beetle
820146
Pyrrhoclerus platyocephalus n. gen. n. sp. Zhang 1989 checkered beetle
820217
Plastophagus pellous n. gen. n. sp. Zhang 1989 checkered beetle
820052
 Coleoptera - Tenebrionidae
Allecula austriaca n. sp. Zhang 1989 comb-clawed beetle
850021
Sinocistela gymnoelytra n. sp. Zhang 1989 comb-clawed beetle
820140
Sinocistela siphla Zhang 1989 comb-clawed beetle
820275
Elytrophagus porphyroleucus n. gen. n. sp. Zhang 1989 darkling beetle
840065
 Coleoptera - Helotidae
"Tarrodacne palmus n. gen. n. sp." = Helota palmus
"Tarrodacne palmus n. gen. n. sp." = Helota palmus Zhang 1989 beetle
830152
 Coleoptera - Carabidae
Carabus mecothoracus n. sp. Zhang 1989 ground beetle
840013
Calosoma cf. maderae Fabricius 1775 ground beetle
820260, 840131
Chlaenius furvus Hong 1984 ground beetle
820187
Anisodactylus giganteus n. sp. Zhang 1989 ground beetle
840102
Miolebidia gaura n. gen. n. sp. Zhang 1989 ground beetle
820141
 Dicondylia -
Lepidoptera indet.1 Linnaeus 1758 moth
820157
 Hymenoptera - Braconidae
Doryctes huadongensis n. sp. Zhang 1989 wasp
830092
Bracon laceolata n. sp. Zhang 1989 wasp
840120
 Hymenoptera - Ichneumonidae
Ovipterygium prominulum n. gen. n. sp., Areoleta quadrivenosa n. gen. n. sp.
Ovipterygium prominulum n. gen. n. sp. Zhang 1989 ichneumon wasp
840001
Areoleta quadrivenosa n. gen. n. sp. Zhang 1989 ichneumon wasp
820265
 Hymenoptera - Scoliidae
"Campsomeris prismatica" = Megacampsomeris prismatica, ? Scolia distincta n. sp.
"Campsomeris prismatica" = Megacampsomeris prismatica Smith 1855 wasp
840052
? Scolia distincta n. sp. Zhang 1989 wasp
830037
 Hymenoptera - Ampulicidae
Longivenula linquensis n. gen. n. sp.
Longivenula linquensis n. gen. n. sp. Zhang 1989 cockroach wasp
820144
 Hymenoptera - Apidae
"Apis miocenica" = Apis miocenica, "Apis fota n. sp." = Apis miocenica, "Apis shandongica n. sp." = Apis miocenica
"Apis miocenica" = Apis miocenica Hong 1984 Chinese Miocene honey bee
820153
"Apis fota n. sp." = Apis miocenica Hong 1984 Chinese Miocene honey bee
820036, 820126, 830072
"Apis shandongica n. sp." = Apis miocenica Hong 1984 Chinese Miocene honey bee
820158
 Hymenoptera - Crabronidae
Megapison silvester n. gen. n. sp., Pison glyptum n. sp.
Megapison silvester n. gen. n. sp. Zhang 1989 apoid wasp
820015
Pison glyptum n. sp. Zhang 1989 apoid wasp
820125
 Hymenoptera - Formicidae
Miosolenopsis fossilis n. gen. n. sp., "Aphaenogaster (Deromyrma) paludosa n. sp." = Paraphaenogaster paludosa, "Aphaenogaster (Sinaphaenogaster) lapidescens n. sp." = Paraphaenogaster lapidescens, Camponotus luteus, Camponotus shanwangensis, Camponotus mitis, Camponotus ambon, Camponotus plenus n. sp., Formica "sp. 2", Formica aff. fusca, Formica ovala n. sp., Formica ceps n. sp., Oecophylla xiejiaheensis, Lasius mordicus n. sp., "Tylolasius inflatus n. gen. n. sp." = Lasius inflatus, Liometopum eremicum n. sp., Elaphrodites mutatus n. sp., Dolichoderus lacinius, Dolichoderus jiaoyanshanensis, Iridomyrmex shandongicus n. sp., "Euponera nubeculata n. sp." = Pachycondyla nubeculata, "? Euponera minutansata n. sp." = Pachycondyla minutansata, Leptogenys lacerata n. sp.
Miosolenopsis fossilis n. gen. n. sp. Zhang 1989 ant
820039, 820080
"Aphaenogaster (Deromyrma) paludosa n. sp." = Paraphaenogaster paludosa Zhang 1989 ant
830080
"Aphaenogaster (Sinaphaenogaster) lapidescens n. sp." = Paraphaenogaster lapidescens Zhang 1989 ant
830136
Camponotus luteus Smith 1858 ant
820161, 840092
Camponotus shanwangensis Hong 1984 ant
820186
Camponotus mitis Smith 1892 ant
820212, 830077
Camponotus ambon Zhang 1989 ant
820035
Camponotus plenus n. sp. Zhang 1989 ant
820172, 820121, 820185
Formica "sp. 2" Linnaeus 1758 ant
820250
Formica aff. fusca Linnaeus 1758 ant
830138
Formica ovala n. sp. Zhang 1989 ant
820160, 840077
Formica ceps n. sp. Zhang 1989 ant
820020
Oecophylla xiejiaheensis Hong 1984 weaver ant
820041
Lasius mordicus n. sp. Zhang 1989 ant
820143
"Tylolasius inflatus n. gen. n. sp." = Lasius inflatus Zhang 1989 ant
820182
Liometopum eremicum n. sp. Zhang 1989 ant
820170
Elaphrodites mutatus n. sp. Zhang 1989 ant
820094
Dolichoderus lacinius Zhang 1989 ant
830075, 840113
Dolichoderus jiaoyanshanensis Hong 1985 ant
840088
Iridomyrmex shandongicus n. sp. Zhang 1989 ant
830127
"Euponera nubeculata n. sp." = Pachycondyla nubeculata Zhang 1989 ant
820065, 820100
"? Euponera minutansata n. sp." = Pachycondyla minutansata Zhang 1989 ant
820115
Leptogenys lacerata n. sp. Zhang 1989 ant
820055
 Hymenoptera - Thynnidae
Brachymethoca gigantea n. gen. n. sp.
Brachymethoca gigantea n. gen. n. sp. Zhang 1989 wasp
820257
 Hymenoptera - Tiphiidae
Tiphia dolichogaster n. sp. Zhang 1989 wasp
820070
 Hymenoptera - Vespidae
Vespa sp. Linnaeus 1758 wasp
840030
Vespa binghami du Buysson 1904 wasp
820003
Rotundipetiolus longiantennus n. gen. n. sp. Zhang 1989 potter wasp
820268
 Hymenoptera - Xyelidae
Megaxyela yaoshanica n. sp., "Xyela cenozoica n. sp." = Enneoxyela cenozoica
Megaxyela yaoshanica n. sp. Zhang 1989 sawfly
840097
"Xyela cenozoica n. sp." = Enneoxyela cenozoica Zhang 1989 sawfly
820138
 Hymenoptera - Cimbicidae
Cimbex chromoptera n. sp., "Clavellaria bicolor n. sp." = Pseudoclavellaria bicolor, "? Clavellaria autochthna n. sp." = Pseudoclavellaria autochthona, "Clavellaria longiclava" = Pseudoclavellaria longiclava, Abia maculosa n. sp.
Cimbex chromoptera n. sp. Zhang 1989 cimbicid sawfly
820017
"Clavellaria bicolor n. sp." = Pseudoclavellaria bicolor Zhang 1989 cimbicid sawfly
820096
"? Clavellaria autochthna n. sp." = Pseudoclavellaria autochthona Zhang 1989 cimbicid sawfly
830045
"Clavellaria longiclava" = Pseudoclavellaria longiclava Zhang 1989 cimbicid sawfly
840006
Abia maculosa n. sp. Zhang 1989 cimbicid sawfly
820091
 Hymenoptera - Tenthredinidae
? Selandria plagineura n. sp. Zhang 1989 wood wasp
830094
Selandria magica n. sp. Zhang 1989 wood wasp
820111
Tenthredo rhysia n. sp. Zhang 1989 wood wasp
830106
Tenthredoides plurilocellata n. gen. n. sp. Zhang 1989 wood wasp
830156
 Hemiptera - Nepidae
Laccotrephes cf. robustus Stal 1870 true bug
830099
 Hemiptera - Corixidae
Corixa parvoculata n. sp. Zhang 1989 water boatman
820131
 Hemiptera - Coreidae
Pseudodasynus nativus n. gen. n. sp.
Pseudodasynus nativus n. gen. n. sp. Zhang 1989 leaf-footed bug
820104
 Hemiptera - Pentatomidae
Necanicarbula shanwangiana n. gen. n. sp. Zhang 1989 stink bug
820061
 Hemiptera - Urostylididae
Urochela nigilimbosa n. sp. Zhang 1989 true bug
820149
Urochela palilionacea n. sp. Zhang 1989 true bug
830064
 Hemiptera - Fulgoridae
Limois pardalis n. sp. Zhang 1989 fulgorid planthopper
830135
Limois shanwangensis Hong 1979 fulgorid planthopper
840119
Aphana lithoecia Zhang 1989 fulgorid planthopper
830085
 Odonata - Macromiidae
Macromia sp. Rambur 1842 river cruiser
820023, 820180, 820183, 840056
 Odonata - Gomphidae
Miopetalura shanwangica n. sp. Zhang 1989 clubtail dragonfly
820013
 Odonata - Aeshnidae
"Mediaeschna lucida" = Epiaeschna lucida, "Aeschna ignivora n. sp." = Aeshna ignivora
"Mediaeschna lucida" = Epiaeschna lucida Zhang 1989 darner dragonfly
820179
"Aeschna ignivora n. sp." = Aeshna ignivora Zhang 1989 hawker dragonfly
820297
 Ephemeroptera - Heptageniidae
Ecdyonurus sp. Eaton 1868 stream mayfly
820081
Heptagenia sp. Walsh 1863 stream mayfly
820147
 Diptera - Bibionidae
Bibio tranopterus n. sp. Zhang 1989 march fly
820155
Bibio eximius n. sp.2 Zhang et al. 1994 march fly
830100
Bibio longineurus n. sp. Zhang 1989 march fly
820133
Bibio peraffinis n. sp. Zhang 1989 march fly
820103
Bibio varineurus n. sp. Zhang 1989 march fly
820078
Bibio cratopodus n. sp. Zhang 1989 march fly
820112
Bibio brachypodus n. sp. Zhang 1989 march fly
820227
Bibio shanwangensis Hong 1984 march fly
820218, 820233
Bibio brunneifemus n. sp. Zhang 1989 march fly
820173
Plecia diatoma n. sp. Zhang 1989 love bug
830017
Plecia vulcania n. sp. Zhang 1989 love bug
840058
Plecia rectivenia n. sp. Zhang 1989 love bug
820139
Plecia villosa Zhang 1989 love bug
840096
 Diptera - Sciaridae
Sciara atropha n. sp. Zhang 1989 dark-winged fungus gnat
840112
 Diptera - Chironomidae
Clinotanypus vagans n. sp. Zhang 1989 non biting midge
830097
 Diptera - Dryomyzidae
Dryomyza shanwangensis n. sp. Zhang 1989 fly
840116