El Simbolar (upper Lumbrera) (Eocene of Argentina)

Also known as above the “Faja Verde II”

Where: Salta, Argentina (25.7° S, 65.4° W: paleocoordinates 28.1° S, 56.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Casamayoran mammal zone, Upper Member (Lumbrera Formation), Middle Eocene (47.8 - 37.7 Ma)

• Lumbrera Formation, above the “Faja Verde II” (Upper Lumbrera)

•Powell et al., 2011: Recently, geologic and paleontologic evidence allowed the correlation of the upper levels of Lumbrera with Geste, Casa Grande, and Quebrada de los Colorados Formations. Additionally, these levels were referred to the late-middle Eocene (Casamayoran SALMA -South American Land Mammal Age-, Barrancan subage) by an U/Pb dating of 39.9 Ma . The lower levels of the Lumbrera Formation are here referred to the Vacan subage of the Casamayoran SALMA.

•Fernicola et al. 2021 obtained an 238U-206Pb isochron age of 46.2 +/- 7.3 Ma for samples from the Lower Lumberer Fm, suggesting that the Lower section of the formation was deposited between 55–46Ma. "According to different authors, the fossiliferous levels of the lower section of the Lumbrera Fm. are equivalent to the Vacan Subage (del Papa et al. 2010; Powell et al. 2011; García-López et al. 2019). However, this biochronology is not supported by the absolute dating provided in this work. The top of the LSLF is now established by radioisotopy at 46. 2 ± 7.3 Ma (early Lutetian), and the record of the EECO and ETM2 in the first 150 m of the LSLF set the initial deposition of the sec- tion in the early Eocene (early Ypresian). With these new constraints, the species based on specimens recovered on the lower levels of the LSLF (this work) are chronologically equivalent to the Itaboraian SALMA (56.1–51.4 Ma; Woodburne et al. 2014; Krause et al. 2017)."

•The age of the Upper Lumbrera Formation has been estimated in 39.9 +/- 0.4 Ma (U/Pb zircon recovered from a tuff layer close to the upper contact, see del Papa et al., 2010). The age of the fossil is therefore considered as middle Eocene (Bartonian)

• bed-level stratigraphic resolution

Environment/lithology: lacustrine; green mudstone and red siltstone

• The paleoenvironmental reconstruction suggests that the Upper Lumbrera Formation represents sedimentation in a fine-grained plain, with localized fluvial and ephemeral lacustrine deposits (del Papa, 2006). Indeed, based on granulometric and petrographic analysis of the Upper Lumbrera sediments at El Simbolar, Lapiana et al. (2016) suggested a paleoenvironment of open plains constructed by the aggradation of reworked loess.
• thick succession of reddish siltstones and sandy siltstones, with sporadic and localized intercalations of fine-grained sandstones and nodules of gypsum/anhydrite. The most outstanding features of this geological unit are its homogeneous lithology and the presence of conspicuous paleosol layers (del Papa et al., 2017).

Size class: macrofossils

Primary reference: D. A. García López and J. E. Powell. 2009. Un nuevo Oldfieldthomasiidae (Mammalia: Notoungulata) del Paleógeno de la provincia de Salta, Argentina. Ameghiniana 46(1):153-164 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 132937: authorized by Philip Mannion, entered by Philip Mannion on 29.08.2012, edited by Richard Butler, Jelle Zijlstra and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Loricata - Sebecidae
cf. Sebecus sp.4 Stirton 1953 crocodilian
PVL 6386, almost complete articulated skull and mandibles, and three cervical vertebrae, exceptionally preserved.
Mammalia
 Cingulata - Dasypodidae
? Utaetus magnum n. sp.3 Herrera et al. 2021 armadillo
Holotype: IBIGEO-P 109, 11 fixed, two movables, and eight caudal complete osteoderms, in addition to numerous fragmentary osteoderms, all from the same individual
 Notoungulata - Campanorcidae
Campanorco sp.4 Bond et al. 1984 notoungulate
“Campanorco inauguralis”; PVL 6225, almost complete skull and upper dentition; IBIGEO-P 105
 Notoungulata - Oldfieldthomasiidae
Dolichostylodon saltensis n. gen. n. sp. García López and Powell 2009 notoungulate
PVL 6219 - holotype (partial skull)
 Notoungulata - Leontiniidae
cf. Coquenia sp.4, Coquenia bondi n. gen. n. sp.1
cf. Coquenia sp.4 Deraco et al. 2008 notoungulate
PVL 6397, articulated specimen with complete skull and jaws, and the anterior third postcra- nial skeleton with almost complete forelimbs.
Coquenia bondi n. gen. n. sp.1 Deraco et al. 2008 notoungulate
PVL 5853 (holotype). PVL 5854
 Notoungulata - Notohippidae
Pampahippus secundus2 Deraco and GarcÍa-López 2016 notoungulate
IBIGEO-P 101