Hainin well vertebrate fauna (Paleocene of Belgium)

Also known as Mons Basin

Where: Hainaut, Belgium (50.4° N, 3.8° E: paleocoordinates 43.8° N, 2.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Hainin Formation, Danian (66.0 - 61.6 Ma)

• Reference-level MP1-5 of the mammalian biochronological scale for the European Palaeogene.

•Although not precisely dated, has been correlated with the earliest Selandian based on stratigraphy (Steurbaut 1998), and more recently to the latest Danian (Steurbaut & Sztrakos 2008).

•The borehole that yielded the fossils encountered continental deposits between 8.50 m and 38.52 m in depth. Of the 30 m of continental deposits, only the section between 11 and 25 m yielded vertebrates (De Bast and Smith 2016).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; claystone

Size classes: macrofossils, mesofossils

Collected by Geological laboratory of the Faculté Polytechnique de Mons in the 1970s

Collection methods: core, chemical

• The sediments collected from the Hainin well went through a first step of screen washing. The remaining hard parts were treated with formic acid (5%) with tricalcic phosphate (Ca3(PO4)2) used as a buffer. A second step of screenwashing was then performed.

Primary reference: E. De Bast, B. Sigé, and T. Smith. 2012. Diversity of the adapisoriculid mammals from the early Palaeocene of Hainin, Belgium. Acta Palaeontologica Polonica 57(1):35-52 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 132949: authorized by Philip Mannion, entered by Philip Mannion on 30.08.2012, edited by Terri Cleary and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

Osteichthyes
 Salientia -
Anura indet. Fischer von Waldheim 1813 frog
Different species of anurans and urodeles (Folie 2007 [SVP abstract])
 Caudata - Urodela
Urodela indet. Duméril 1805 salamander
Different species of anurans and urodeles (Folie 2007 [SVP abstract])
Reptilia
 Testudines - Trionychidae
"? Palaeotrionyx sp." = Paleotrionyx6, Trionyx sp.6
"? Palaeotrionyx sp." = Paleotrionyx6 Schmidt 1945 softshell turtle
costal fragments
Trionyx sp.6 Geoffroy Saint-Hilaire 1809 softshell turtle
carapace fragments
 Testudines -
cf. Baenoidea indet.6 turtle
shell fragments
 Squamata - Lacertidae
Lacertidae indet.1 Bonaparte 1831 lizard
 Squamata - Anguidae
Anguini indet.1 Augé 2005 lizard
dentary fragments
 Squamata -
Alethinophidia indet.7 Nopcsa 1923 snake
Three fragmentary vertebrae and one fragmentary rib; two referred to cf. Dunnophis by Van Dyck (1983)
 Squamata - Amphisbaenidae
Amphisbaenidae indet.1 Gray 1865 worm lizard
H13, 70-14, 01; H14-14, 60, 03; H14, 60-15, 10, 03; H14, 60-15, 10, 01: four dorsal vertebrae
 Squamata - Polyodontobaenidae
Polyodontobaena belgica n. gen. n. sp.5
Polyodontobaena belgica n. gen. n. sp.5 Folie et al. 2013 worm lizard
IRSNB R 283 (HA 605) - holotype (nearly complete left dentary); paratype: IRSNB R 284 (HA 606), a right dentary; referred material: 13.7-14 (01), 16.4-15.10 (01) and HA 607- 639 (37 trunk vertebrae including HA 607/IRSNB R 285); HA 640-642 (3 cervical or caudal vertebrae including HA 640/IRSNB R 286)
 Squamata -
Scincoideus haininensis n. gen. n. sp.4
Scincoideus haininensis n. gen. n. sp.4 Folie et al. 2005 lizard
IRSNB R 219 - holotype (right dentary); paratypes: four maxillae and maxillary fragments (IRSNB R 215-218), eight dentaries and dentary fragments (IRSNB R 220-227), one pair of frontals (IRSNB R 228), two isolated frontals (IRSNB R 229-230) and one incomplete parietal (IRSNB R 231)
Mammalia
 Eutheria -
Cimolesta indet.3 eutherian
IRSNB M2225
 Cimolesta - Cimolestidae
cf. Cimolestes sp.3 Marsh 1889 eutherian
IRSNB M2198
 Cimolesta - Palaeoryctidae
Belgoryctes thaleri n. gen. n. sp.3
Belgoryctes thaleri n. gen. n. sp.3 De Bast and Smith 2017 eutherian
IRSNB M2193 - holotype; referred material: IRSNB M1947 and IRSNB M2194-2197
 Cimolesta - Pentacodontidae
Eurolestes dupuisi n. gen. n. sp.3
Eurolestes dupuisi n. gen. n. sp.3 De Bast and Smith 2017 eutherian
IRSNB M2203 - holotype; referred material: IRSNB M1945 and 2199-2205
 Primates - Picrodontidae
Russellodon haininense n. gen. n. sp.3
Russellodon haininense n. gen. n. sp.3 Sigé and Marandat 1997 primate
IRSNB M1977 - holotype; referred material: IRSNB M2211 and 2212
 Tribosphenida - Adapisoriculidae
Afrodon gheerbranti n. sp., cf. Adapisoriculus sp.3, "Peradectes marandati n. sp." = Bustylus marandati2, Bustylus folieae n. sp., Proremiculus lagnauxi n. gen. n. sp.
Afrodon gheerbranti n. sp. De Bast et al. 2012 placental
IRSNB M1982 - type (right M2); paratypes: IRSNB M1982-1989
cf. Adapisoriculus sp.3 Lemoine 1885 placental
IRSNB M2009 (right M3)
"Peradectes marandati n. sp." = Bustylus marandati2 Crochet and Sigé 1983 placental
IRSNB M1974 - type (right M2); and numerous referred material
Bustylus folieae n. sp. De Bast et al. 2012 placental
IRSNB M1996 - type (left M1); and numerous referred material
Proremiculus lagnauxi n. gen. n. sp. De Bast et al. 2012 placental
IRSNB M2006 - holotype (left M2); and numerous referred material
 Condylarthra - Pleuraspidotheriidae
? Pleuraspidotheriidae indet.3 Zittel 1892 placental
IRSNB M2222
 Tribosphenida -
Prolatidens waudruae n. gen. n. sp.8, Quadratodon sigei n. gen. n. sp.3
Prolatidens waudruae n. gen. n. sp.8 Sudre and Russell 1982 placental
IRSNB M1972 - holotype (left M1); referred material: IRSNB M2053 (= N2-15), left P4; IRSNB M2054 (= Q2-19), leftM1 orM2; IRSNBM2055 (= Q1-16), right M1 or M2; IRSNB M2056 (= O1-05), left dp4; IRSNB M2115 (= MN1.2-03), left m1; IRSNB M2116 (= N2-01), right m1; IRSNB M1973 (= O1-01), left m2; IRSNB M2117 (= N1-05), right m1 or m2 (De Bast and Smith 2013)
Quadratodon sigei n. gen. n. sp.3 De Bast and Smith 2017 placental
IRSNB M2221 - holotype; paratypes: IRSNB M2218 and 2220; referred material: IRSNB M2216-2219
 Tribosphenida - Pseudorhyncocyonidae
Pseudorhyncocyonidae indet.3 Sigé 1974 placental
IRSNB M2224
 Macroscelidea - Louisinidae
Monshyus praevius n. gen. n. sp.8, Cingulodon magioncaldai n. gen. n. sp.3
Monshyus praevius n. gen. n. sp.8 Sudre and Russell 1982 elephant shrew
IRSNB M1970 - holotype (left M1 or M2); referred material: R1-7 (right DP4)
Cingulodon magioncaldai n. gen. n. sp.3 De Bast and Smith 2017 elephant shrew
IRSNB M2223 - holotype
 Multituberculata - Kogaionidae
Hainina belgica n. gen. n. sp.9, Hainina godfriauxi n. sp.9
Hainina belgica n. gen. n. sp.9 Vianey-Liaud 1979 multituberculate
IRSNB M1964 - holotype; referred material: IRSNB M1956-1963, 2185-2190
Hainina godfriauxi n. sp.9 Vianey-Liaud 1979 multituberculate
IRSNB M1968 - type; referred material: IRSNB M1955, 1965-1969 and 2191-2192
 Multituberculata - Boffiidae
Boffius splendidus n. gen. n. sp.9
Boffius splendidus n. gen. n. sp.9 Vianey-Liaud 1979 multituberculate
IRSNB M1948 - holotype; referred material: IRSNB M1949-1954