PBDB Collection

La Barda, Paso del Sapo (Eocene of Argentina)

Also known as Bardalestes hunco type - Estancia 26 de Mayo

Where: Chubut, Argentina (42.8° S, 69.9° W: paleocoordinates 45.3° S, 60.1° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Andesitas Huancache Formation, Lutetian (47.8 - 41.3 Ma)

• These include a fauna from the Tufolitas Laguna del Hunco Formation

•at Laguna Fría (42° 43’ 31.5” S, 69° 51’ 36.1” W) and another from the interbedded tuffs of the

•Andesitas Huancache Formation at the La Barda locality (42° 46’ 48.5” S, 69° 51’ 43.3”) (Fig. 1). The

•age of these faunas is constrained by radioisotopic dates from nearby volcanic deposits which underlie

•and overlie the mammal-bearing levels. The Laguna Fría fauna unconformably overlies a basal

•ignimbrite dated by 40Ar-39Ar method at 49.51 ± 0.32 and it underlies the Andesitas Huancache alkaline

•basalt dated at 47.89 ± 1.21 Ma (Tejedor et al., 2009). At the second locality, the La Barda Tuffs are

•interbedded with the Andesitas Huancache lava flows, and they occur above a basalt dated by the 40K40Ar method to 47.89 ± 1.21 Ma (Tejedor et al., 2009). This indicates that the Paso del Sapo faunas are

•late early Eocene in age (Ypresian-Lutetian boundary).

• bed-level stratigraphic resolution

Environment/lithology: fluvial; tuff

• Lorente et al 2016: Mammal-bearing levels at the La Barda locality occur within the Andesitas Huancanche interbedded tuffs. The Andesitas Huancanche are part of the so-called Middle Chubut River Volcanic–Pyroclastic Complex, the geology and stratigraphy of which was reviewed by Arag?on & Mazzoni (1997). Additional information on the regional geology, stratigraphy, geochemistry, petrogenesis and palaeobotany of this complex has been provided by Peter- sen (1946), Archangelsky (1974), Volkheimer & Lage (1981), Lage (1982), Rapela et al. (1984), Arag?on & Romero (1984), Arag?on et al. (1987), Mazzoni & Arag?on (1985) and Mazzoni et al. (1989). The Andesitas Huan- cache Upper Member comprises alkali basalts dated of these levels between 45 and 47 Ma was proposed by between 47.89 ? 1.21and 43 Ma. A time of deposition Tejedor et al. (2009).

Size class: mesofossils

Collected in 2005–2010

Collection methods: bulk, surface (float), sieve,

Primary reference: F. J. Goin, A. M. Candela, M. A. Abello and E. V. Oliveira. 2009. Earliest South American paucituberculatans and their significance in the understanding of ‘pseudodiprotodont’ marsupial radiations. Zoological Journal of the Linnean Society 155:867-884 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 133039: authorized by Philip Mannion, entered by Philip Mannion on 31.08.2012, edited by Matthew Carrano, Jelle Zijlstra and Miranta Kouvari

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Mammalia
Tribosphenida -	Marmosopsis sp.⁴, Palangania sp.⁴, Palangania cf. brandmayri⁴ Marmosopsis sp.⁴ Paula Couto 1962 metatherian two right mx (LIEB-PV 1286, and LIEB-PV 1287) Palangania sp.⁴ Goin et al. 1998 marsupial LIEB-PV 1106, left M?3; fig. 3J Palangania cf. brandmayri⁴ Goin et al. 1998 marsupial Two lower molars from La Barda (LIEB- PV 1089 and 1090)
Marsupialia -	Paucituberculata "gen. et sp. nov. 5"⁴, ? Paucituberculata "gen. et sp. indet. B"⁴ Paucituberculata "gen. et sp. nov. 5"⁴ Ameghino 1894 marsupial one upper (LIEB-PV 1151) and three lower molars (LIEB-PV 1101, 1136) ? Paucituberculata "gen. et sp. indet. B"⁴ Ameghino 1894 marsupial left mandible
Paucituberculata -	Bardalestes hunco n. gen. n. sp. Bardalestes hunco n. gen. n. sp. Goin et al. 2009 marsupial LIEB-PV 1135 - holotype (a fragment of left maxillary with M2–3, and roots and part of the crown of M4)
Tribosphenida - Peradectidae	"Caroloameghiniidae "gen. et sp. nov. 2"" = Caroloameghiniinae⁴, "? Caroloameghiniidae indet." = Caroloameghiniinae⁴ "Caroloameghiniidae "gen. et sp. nov. 2"" = Caroloameghiniinae⁴ Ameghino 1901 marsupial one upper molar from La Barda (LIEB-PV 1098, a left MX) "? Caroloameghiniidae indet." = Caroloameghiniinae⁴ Ameghino 1901 marsupial isolated, left mx talonid (LIEB- PV 1099)
Tribosphenida -	Gashternia ctalehor⁴ Gashternia ctalehor⁴ Simpson 1935 marsupial Several upper and lower molars
Marsupialia -	Didelphimorphia "gen. et sp. nov. 3"⁴ Didelphimorphia "gen. et sp. nov. 3"⁴ Gill 1872 marsupial Several upper and lower molars
Tribosphenida - Protodidelphidae	Protodidelphis "sp. nov."⁴, Protodidelphis sp.⁴ Protodidelphis "sp. nov."⁴ Paula Couto 1952 marsupial several lower and upper isolated teeth, including LIEB-PV 1097, left M3 Protodidelphis sp.⁴ Paula Couto 1952 marsupial Two isolated lower molars (LIEB-PV 1092, isolated left m?1; LIEB-PV 1142, isolated left m?3)
Tribosphenida - Derorhynchidae	Pauladelphys sp.⁴ Pauladelphys sp.⁴ Goin et al. 1999 marsupial a few specimens
Tribosphenida -	Australidelphia indet.³ Australidelphia indet.³ Szalay 1982 marsupial one astragalus and three calcanea
Marsupialia -	? Microbiotheria "gen. et sp. indet. F"⁴ ? Microbiotheria "gen. et sp. indet. F"⁴ Ameghino 1899 marsupial two upper molars
Microbiotheria - Microbiotheriidae	? Microbiotheriidae "gen. et sp. indet. E"⁴ ? Microbiotheriidae "gen. et sp. indet. E"⁴ Ameghino 1887 marsupial LIEB-PV 1094 is an upper right molar lacking the protocone, probably an M4
Polydolopimorphia - Polydolopidae	Polydolopidae indet.¹, "Polydolops rothi" = Pliodolops rothi⁴, Polydolops unicus n. sp.⁴, Amphidolops "sp. nov. 2"⁴, Amphidolops intermedius n. sp.¹, Hypodolops sapoensis¹, Hypodolops sapoensis¹ Polydolopidae indet.¹ Ameghino 1897 metatherian LIEB-PV 1178 is a right mandibular fragment with p3–m1 "Polydolops rothi" = Pliodolops rothi⁴ Simpson 1936 metatherian several upper and lower specimens Polydolops unicus n. sp.⁴ Tejedor et al. 2009 metatherian Holotype, LIEB-PV 1252, right maxillary fragment with P3–M3; LIEB-PV 1245, left maxillary fragment with M1–2; LIEB-PV 1246, left M1; LIEB-PV 1247, left M2; LIEB-PV 1248, left M2; LIEB-PV 1249, left M2; LIEB-PV 1250, left M2; LIEB-PV 1251, right maxillary fragment with P3–M3; LIEB-PV 1253, right M1; LIEB-PV 1254, right M1; LIEB-PV 1255, right M1; LIEB- PV 1256, right, broken M2; LIEB-PV 1257, right M3; LIEB-PV 1258, right M3; LIEB-PV 1263, left mandible with m1–2; LIEB-PV 1265, left m2; LIEB-PV 1266, left, broken m2; LIEB-PV 1267, right mandible with broken p3, complete m1–2, and roots of m3; LIEB-PV 1268, right mandible with m2; LIEB-PV 1269, right mandible with m2; LIEB-PV 1270, right mandible with m2; LIEB-PV 1271, right mandible with broken m1, complete m2, and roots of m3; LIEB-PV 1272, left m1; LIEB-PV 1273, right m1; LIEB-PV 1274, right m1; LIEB-PV 1275, right m2; LIEB-PV 1276, broken right m2; LIEB-PV 1277, right m3; LIEB-PV 1278, right m3. Amphidolops "sp. nov. 2"⁴ Ameghino 1902 metatherian Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1187, a right m2, and several isolated lower and upper molars from both localities of Paso del Sapo" Amphidolops intermedius n. sp.¹ Chornogubsky 2020 metatherian type: LIEB-PV 1187, LIEB-PV 1188, LIEB-PV 1181, LIEB-PV 1182, LIEB-PV 1183, LIEB-PV 1184, LIEB-PV 1185 and LIEB-PV 1186 Hypodolops sapoensis¹ Chornogubsky 2020 metatherian Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1153, a maxillary fragment with M1–2, as well as four lower molars are included in this new species recovered in La Barda and Laguna Fría" Hypodolops sapoensis¹ Chornogubsky 2020 metatherian Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1173, a right maxillary fragment with M1–2, as well as several maxillary and mandibular remains and isolated teeth have been assigned to this new taxon collected in both localities of Paso del Sapo."
Sparassodonta - Hathliacynidae	"Hathliacynidae "gen. et sp. nov. 6"" = Hathlyacininae⁴ "Hathliacynidae "gen. et sp. nov. 6"" = Hathlyacininae⁴ Ameghino 1894 metatherian LIEB-PV 1141, a fragment of a left M?2
Sparassodonta - Borhyaenidae	Borhyaeninae "gen. et sp. indet. C"⁴ Borhyaeninae "gen. et sp. indet. C"⁴ Ameghino 1894 metatherian LIEB-PV 1093 is a right m?1
Panameriungulata - Protolipternidae	Asmithwoodwardia "sp. nov."⁴, Asmithwoodwardia subtrigona⁴ Asmithwoodwardia "sp. nov."⁴ Ameghino 1901 placental m1 or m2 (LIEB-PV 1613) Asmithwoodwardia subtrigona⁴ Ameghino 1901 placental
Cingulata - Dasypodidae	Prostegotherium cf. astrifer⁴, Riostegotherium sp.⁴, Stegosimpsonia sp.⁴ Prostegotherium cf. astrifer⁴ Ameghino 1902 armadillo osteoderms Riostegotherium sp.⁴ Oliveira and Bergqvist 1998 armadillo Several isolated plates of the dorsal carapace Stegosimpsonia sp.⁴ Vizcaino 1994 armadillo several isolated plates of the dorsal carapace
Notoungulata - Archaeopithecidae	Archaeopithecus cf. rogeri⁴ Archaeopithecus cf. rogeri⁴ Ameghino 1897 notoungulate Not clear which material comes from Laguna Fría and which from La Barda. "Several maxillary and mandibular fragments (e.g., LIEB-PV 1625 to 1628), as well as isolated teeth (e.g., LIEB-PV 1629) referred to this family have been found in La Barda and Laguna Fría."
Notoungulata - Henricosborniidae	Henricosbornia lophodonta⁴ Henricosbornia lophodonta⁴ Ameghino 1901 notoungulate "Maxillary and mandibular fragments re- ferred to the Family Henricosborniidae are abundant in both localities of Paso del Sapo, with complete series of premolars and molars, as well as isolated teeth."
Notoungulata - Notostylopidae	Notostylops sp.⁴, Homalostylops parvus⁴ Notostylops sp.⁴ Ameghino 1897 notoungulate Mandibular fragments and isolated upper teeth that can be referred to the genus Notostylops are abundant in both localities of Paso del Sapo Homalostylops parvus⁴ Ameghino 1897 notoungulate mandibular fragment with m1–2 of a small notostylopid was collected in La Barda (LIEB-PV 1621).
Allotheria - Sudamericidae	Greniodon sylvaticus n. gen. n. sp.² Greniodon sylvaticus n. gen. n. sp.² Goin et al. 2012 mammal LIEB-PV 2000, 2001

¹Chornogubsky 2020; ²Goin et al. 2012; ³Lorente et al. 2016; ⁴Tejedor et al. 2009