Sample A, Titusville Site (1967 excavation) (Pleistocene of the United States)

Where: Pennsylvania (41.6° N, 79.6° W: paleocoordinates 41.6° N, 79.6° W)

• coordinate stated in text

• hand sample-level geographic resolution

When: Titusville Till Formation, Late/Upper Pleistocene (0.1 - 0.0 Ma)

Environment/lithology: mire or swamp; peat

• Like Zone I, analogues to the Zone III fauna occur today in forest–tundra environments of North America where mean July temperatures range from 10 to 13C.

Size class: mesofossils

Collected in 1967

Collection methods: quarrying, sieve

• All specimens, including those examined by Totten and Coope in 1967, were mounted onto micropaleontological slides and reposited in the Quaternary Entomology Laboratory at North Dakota State University.

Primary reference: S. Cong, A. C. Ashworth, D. P. Schwert and S. M. Totten. 1996. Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania. Quaternary Research 45:216-225 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 137209: authorized by Matthew Clapham, entered by Jered Karr on 12.12.2012, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Coleoptera - Staphylinidae
"Holoboreaphilus nordenskioeldi" = Holoboreaphilus nordenskioldi Maklin 1878 rove beetle
Acidota quadrata Zetterstedt 1837 rove beetle
Olophrum rotundicolle Sahlberg 1817 rove beetle
Olophrum consimile Gyllenhal 1810 rove beetle
Pycnoglypta sp. Thomson 1858 winter rove beetle
Pycnoglypta aptera or Pycnoglypta lurida
Pycnoglypta aptera Campbell 1983 winter rove beetle
 Coleoptera - Helophoridae
Helophorus tuberculatus Gyllenhal 1808 water scavenger beetle
 Coleoptera - Carabidae
Dyschirius melancholicus Putzeys 1867 ground beetle
"Pterostichus patruelis" = Pterostichus (Phonias) patruelis Dejean 1831 ground beetle
"Carabus maeander" = Carabus (Homoeocarabus) maeander Fischer von Waldheim 1822 ground beetle
Elaphrus lapponicus Gyllenhal 1810 ground beetle
"Trechus crassiscapus" = Trechus (Trechus) crassiscapus Lindroth 1955 ground beetle
"Bembidion fortestriatum" = Bembidion (Trepanedoris) fortestriatum Motschulsky 1845 ground beetle
Patrobus septentrionis Dejean 1828 ground beetle