Khulsan (general) [PMPE] (Cretaceous of Mongolia)

Also known as Polish-Mongolian Palaeontological Expedition, East Nemegt

Where: Omnogov, Mongolia (43.5° N, 101.1° E: paleocoordinates 40.7° N, 90.7° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Barun Goyot Formation, Campanian (83.6 - 72.1 Ma)

• unspecified stratigraphic information

• group of beds-level stratigraphic resolution

Environment/lithology: eolian; sandstone and pebbly, conglomeratic sandstone

• "The sandstone beds are massive, horizontally stratified or cross-stratified."

Size class: macrofossils

Preservation: mold/impression

Collected by PMPE in 1970-1971

Collection methods: quarrying, surface (float), surface (in situ), mechanical,

Primary reference: Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 13776: authorized by Matthew Carrano, entered by Matthew Carrano on 25.04.2002, edited by Roger Benson, Jonathan Tennant and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Myriapoda
 Spirobolida -
Gobiulus sabulosus n. gen. n. sp.7
Gobiulus sabulosus n. gen. n. sp.7 Dzik 1975 millipede
Aves
  -
Aves indet.14 Linnaeus 1758 bird
eggs; ZPAL MgOv-I/19, I/21a-c, I/25c-d
 Enantiornithes -
Gobipteryx minuta n. gen. n. sp.8
Gobipteryx minuta n. gen. n. sp.8 Elzanowski 1974 bird
Reptilia
 Ornithischia - Ankylosauridae
Ankylosauridae indet. Brown 1908 ornithischian
sull wiith skeleton
 Neornithischia - Protoceratopsidae
Protoceratopsidae indet.14 Granger and Gregory 1923 ornithischian
ZPAL MgOv-I/5-7, 8a-b, 9a-e, 10a, 12-13, 14a-b, 15a-d, 17, 20, 23-24, 26a-f,; P2, P3 group eggs
? Protoceratopsidae indet.14 Granger and Gregory 1923 ornithischian
ZPAL MgOv-I/10, I/25a-b
 Neornithischia -
"? Protoceratops kozlowskii n. sp." = Bagaceratops rozhdestvenskyi12
"? Protoceratops kozlowskii n. sp." = Bagaceratops rozhdestvenskyi12 Maryanska and Osmólska 1975 ornithischian
ZPAL MgD 1/117
 Neornithischia - Bagaceratopsidae
Lamaceratops tereschenkoi n. gen. n. sp.1
Lamaceratops tereschenkoi n. gen. n. sp.1 Alifanov 2003 ornithischian
 Squamata - Paravaranidae
Paravaranus angustifrons n. gen. n. sp.3
Paravaranus angustifrons n. gen. n. sp.3 Borsuk-Bialynicka 1984 lizard
ZPAL MgR I/67 (holotype), damaged skull
 Squamata - Necrosauridae
Proplatynotia longirostrata n. gen. n. sp.3 Borsuk-Bialynicka 1984 lizard
ZPAL MgR I/68 (holotype), posteriorly damaged skull and mandible
 Squamata -
Gobiderma pulchra3 Borsuk-Bialynicka 1984 lizard
ZPAL MgR-I/54, skull fragment
 Squamata - Varanidae
Saniwides mongoliensis n. gen. n. sp.3 Borsuk-Bialynicka 1984 monitor lizard
ZPAL MgR-I/72 (holotype), incomplete and deformed skull and mandible
Telmasaurus grangeri3 Gilmore 1943 monitor lizard
ZPAL MgR-I/65, skull lacking preorbital part
 Squamata -
Parviderma inexacta n. gen. n. sp.3
Parviderma inexacta n. gen. n. sp.3 Borsuk-Bialynicka 1984 lizard
ZPAL MgR-I/43 (holotype), very damaged skull with left mandible
 Squamata - Polyglyphanodontidae
Adamisaurus magnidentatus15, "Macrocephalosaurus chulsanensis n. sp." = Gilmoreteius chulsanensis16, "Macrocephalosaurus ? gilmorei" = Gilmoreteius gilmorei12
Adamisaurus magnidentatus15 Sulimski 1972 lizard
ZPAL MgR-I/13 (partial skull, vertebrae and hand), ZPAL MgR-1/35,42 (partial skulls)
"Macrocephalosaurus chulsanensis n. sp." = Gilmoreteius chulsanensis16 Sulimski 1975 lizard
"Macrocephalosaurus ? gilmorei" = Gilmoreteius gilmorei12 Sulimski 1975 lizard
 Squamata -
Polrussia mongoliensis n. gen. n. sp.6
Polrussia mongoliensis n. gen. n. sp.6 Borsuk-Bialynicka and Alifanov 1991 lizard
 Squamata - Phrynosomatidae
Desertiguana gobiensis n. gen. n. sp.2
Desertiguana gobiensis n. gen. n. sp.2 Alifanov 2013 lizard
PIN 4487/0, left mandibular ramus without the occipital end
 Squamata -
Igua minuta n. gen. n. sp.6
Igua minuta n. gen. n. sp.6 Borsuk-Bialynicka and Alifanov 1991 lizard
ZPAL MgR-I/60 (holotype skull)
 Squamata - Priscagamidae
Priscagama gobiensis9 Borsuk-Bialynicka and Moody 1984 lizard
ZPAL MgR-III/69
 Squamata - Slavoiidae
Slavoia darevskii n. gen. n. sp.17
Slavoia darevskii n. gen. n. sp.17 Sulimski 1984 lizard
 Squamata - Carusiidae
Shinisauroides latipalatum4 Borsuk-Bialynicka 1985 lizard
ZPAL MgR-I/56, 55, 57 (partial skulls)
 Squamata -
Globaura venusta5 Borsuk-Bialynicka 1988 lizard
ZPAL MgR-I/45, 46, 48, 49, 51, 55, 118, 47 (partial-complete large skulls), 50, 71 (small skulls)
Eoxanta lacertifrons5 Borsuk-Bialynicka 1988 lizard
ZPAL MgR-I/61, 74 (partial skulls)
Mammalia
 Tribosphenida -
Asioryctes nemegtensis11 Kielan-Jaworowska 1975 eutherian
 Tribosphenida - Zalambdalestidae
Barunlestes butleri n. gen. n. sp.11
Barunlestes butleri n. gen. n. sp.11 Kielan-Jaworowska 1975 eutherian
ZPAL MgM-I/77 (holotype partial skull and postcranium)
 Theriiformes -
Deltatheridium pretrituberculare12 Gregory and Simpson 1926 mammal
subspecies tardum
 Multituberculata - Djadochtatheriidae
"Djadochtatherium catopsaloides" = Catopsbaatar catopsaloides10
"Djadochtatherium catopsaloides" = Catopsbaatar catopsaloides10 Kielan-Jaworowska 1974 multituberculate
fragment of lower jaw
 Multituberculata -
Chulsanbaatar vulgaris n. gen. n. sp.10 Kielan-Jaworowska 1974 multituberculate
36 specimens, including holotype skull (ZPAL MgM-I/139)
Nemegtbaatar gobiensis10 Kielan-Jaworowska 1974 multituberculate
Phaceloolithidae
  - Phaceloolithidae
Dendroolithus sp.13 Zhao and Li 1988
eggs