PBDB Collection

Khulsan (general) [PMPE] (Cretaceous to of Mongolia)

Also known as Polish-Mongolian Palaeontological Expedition, East Nemegt

Where: Omnogov, Mongolia (43.5° N, 101.1° E: paleocoordinates 40.7° N, 90.5° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Baruungoyot Formation, Middle Campanian to Middle Campanian (83.5 - 70.6 Ma)

• unspecified stratigraphic information

• group of beds-level stratigraphic resolution

Environment/lithology: eolian; sandstone and pebbly, conglomeratic sandstone

• "The sandstone beds are massive, horizontally stratified or cross-stratified."

Size class: macrofossils

Preservation: mold/impression

Collected by PMPE in 1970-1971

Collection methods: quarrying, surface (float), surface (in situ), mechanical,

Primary reference: Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 13776: authorized by Matthew Carrano, entered by Matthew Carrano on 25.04.2002, edited by Roger Benson, Jonathan Tennant and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Reptilia
Ornithischia - Ankylosauridae	Ankylosauridae indet. Ankylosauridae indet. Brown 1908 ankylosaurid sull wiith skeleton
Ornithischia - Protoceratopsidae	Protoceratopsidae indet.¹⁴, ? Protoceratopsidae indet.¹⁴, Breviceratops kozlowskii⁶ Protoceratopsidae indet.¹⁴ Granger and Gregory 1923 ceratopsian ZPAL MgOv-I/5-7, 8a-b, 9a-e, 10a, 12-13, 14a-b, 15a-d, 17, 20, 23-24, 26a-f,; P2, P3 group eggs ? Protoceratopsidae indet.¹⁴ Granger and Gregory 1923 ceratopsian ZPAL MgOv-I/10, I/25a-b Breviceratops kozlowskii⁶ Maryanska and Osmólska 1975 ceratopsian
Aves
-	Aves indet.¹⁴ Aves indet.¹⁴ Linnaeus 1758 bird eggs; ZPAL MgOv-I/19, I/21a-c, I/25c-d
Enantiornithes -	Gobipteryx minuta n. gen. n. sp.⁸ Gobipteryx minuta n. gen. n. sp.⁸ Elzanowski 1974 bird
Reptilia
Squamata -	Eoxanta lacertifrons⁴, Globaura venusta⁴ Eoxanta lacertifrons⁴ Borsuk-Bialynicka 1988 squamates ZPAL MgR-I/61, 74 (partial skulls) Globaura venusta⁴ Borsuk-Bialynicka 1988 squamates ZPAL MgR-I/45, 46, 48, 49, 51, 55, 118, 47 (partial-complete large skulls), 50, 71 (small skulls)
Squamata - Carusiidae	Shinisauroides latipalatum³ Shinisauroides latipalatum³ Borsuk-Bialynicka 1985 squamates ZPAL MgR-I/56, 55, 57 (partial skulls)
Squamata - Polyglyphanodontidae	"Macrocephalosaurus ? gilmorei" = Gilmoreteius gilmorei¹², "Macrocephalosaurus chulsanensis n. sp." = Gilmoreteius chulsanensis¹⁶, Adamisaurus magnidentatus¹⁵ "Macrocephalosaurus ? gilmorei" = Gilmoreteius gilmorei¹² Sulimski 1975 squamates "Macrocephalosaurus chulsanensis n. sp." = Gilmoreteius chulsanensis¹⁶ Sulimski 1975 squamates ZPAL MgR-I/14–18, 20, 22–31 Adamisaurus magnidentatus¹⁵ Sulimski 1972 squamates ZPAL MgR-I/13 (partial skull, vertebrae and hand), ZPAL MgR-1/35,42 (partial skulls)
Squamata - Slavoiidae	Slavoia darevskii n. gen. n. sp.¹⁷ Slavoia darevskii n. gen. n. sp.¹⁷ Sulimski 1984 squamates
Squamata - Paravaranidae	Paravaranus angustifrons n. gen. n. sp.² Paravaranus angustifrons n. gen. n. sp.² Borsuk-Bialynicka 1984 squamates ZPAL MgR I/67 (holotype), damaged skull
Squamata - Necrosauridae	Proplatynotia longirostrata n. gen. n. sp.² Proplatynotia longirostrata n. gen. n. sp.² Borsuk-Bialynicka 1984 squamates ZPAL MgR I/68 (holotype), posteriorly damaged skull and mandible
Squamata -	Gobiderma pulchra² Gobiderma pulchra² Borsuk-Bialynicka 1984 squamates ZPAL MgR-I/54, skull fragment
Squamata - Varanidae	Telmasaurus grangeri², Saniwides mongoliensis n. gen. n. sp.² Telmasaurus grangeri² Gilmore 1943 monitor lizard ZPAL MgR-I/65, skull lacking preorbital part Saniwides mongoliensis n. gen. n. sp.² Borsuk-Bialynicka 1984 monitor lizard ZPAL MgR-I/72 (holotype), incomplete and deformed skull and mandible
Squamata -	Parviderma inexacta n. gen. n. sp.², Igua minuta n. gen. n. sp.⁵ Parviderma inexacta n. gen. n. sp.² Borsuk-Bialynicka 1984 squamates ZPAL MgR-I/43 (holotype), very damaged skull with left mandible Igua minuta n. gen. n. sp.⁵ Borsuk-Bialynicka and Alifanov 1991 squamates ZPAL MgR-I/60 (holotype skull)
Squamata - Phrynosomatidae	Desertiguana gobiensis n. gen. n. sp.¹ Desertiguana gobiensis n. gen. n. sp.¹ Alifanov 2013 squamates PIN 4487/0, left mandibular ramus without the occipital end
Squamata -	Polrussia mongoliensis n. gen. n. sp.⁵ Polrussia mongoliensis n. gen. n. sp.⁵ Borsuk-Bialynicka and Alifanov 1991 squamates ZPAL MgR-I/119
Squamata - Priscagamidae	Priscagama gobiensis⁹ Priscagama gobiensis⁹ Borsuk-Bialynicka and Moody 1984 squamates ZPAL MgR-III/69
Mammalia
Multituberculata -	Nemegtbaatar gobiensis¹⁰ Nemegtbaatar gobiensis¹⁰ Kielan-Jaworowska 1974 multituberculate
Multituberculata - Djadochtatheriidae	"Djadochtatherium catopsaloides" = Catopsbaatar catopsaloides¹⁰ "Djadochtatherium catopsaloides" = Catopsbaatar catopsaloides¹⁰ Kielan-Jaworowska 1974 multituberculate fragment of lower jaw
Multituberculata -	Chulsanbaatar vulgaris n. gen. n. sp.¹⁰ Chulsanbaatar vulgaris n. gen. n. sp.¹⁰ Kielan-Jaworowska 1974 multituberculate 36 specimens, including holotype skull (ZPAL MgM-I/139)
Theriamorpha - Zalambdalestidae	Barunlestes butleri n. gen. n. sp.¹¹ Barunlestes butleri n. gen. n. sp.¹¹ Kielan-Jaworowska 1975 eutherian ZPAL MgM-I/77 (holotype partial skull and postcranium)
Theriamorpha - Asioryctidae	Asioryctes nemegtensis¹¹ Asioryctes nemegtensis¹¹ Kielan-Jaworowska 1975 eutherian
Deltatheroida - Deltatheridiidae	Deltatheridium pretrituberculare¹² Deltatheridium pretrituberculare¹² Gregory and Simpson 1926 metatherian subspecies tardum
Myriapoda
Spirobolida -	Gobiulus sabulosus n. gen. n. sp.⁷ Gobiulus sabulosus n. gen. n. sp.⁷ Dzik 1975 millipede
Phaceloolithidae
- Phaceloolithidae	Dendroolithus sp.¹³ Dendroolithus sp.¹³ Zhao and Li 1988 eggs

¹Alifanov 2013; ²Borsuk-Bialynicka 1984; ³Borsuk-Bialynicka 1985; ⁴Borsuk-Bialynicka 1988; ⁵Borsuk-Bialynicka and Alifanov 1991; ⁶Czepiński 2019; ⁷Dzik 1975; ⁸Elzanowski 1974; ⁹Gao and Hou 1995; ¹⁰Kielan-Jaworowska 1974; ¹¹Kielan-Jaworowska 1975; ¹²Maryanska and Osmólska 1975; ¹³Mikhailov et al. 1994; ¹⁴Sabath 1991; ¹⁵Sulimski 1972; ¹⁶Sulimski 1975; ¹⁷Sulimski 1984