PBDB Collection

Shanwang, Linqu County, collection K0 (Shandong Geological Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Geological Museum. This collection contains specimens labeled with three-digit numbers prefaced by K0.

Primary reference: J. F. Zhang. 1989. Miocene insects from Shanwang of Shandong, China and their bearing on palaeoenvironment. Proceedings of International Symposium on Pacific Neogene Continental and Marine Events 149-156 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 138944: authorized by Matthew Clapham, entered by Matthew Clapham on 03.02.2013

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae - Araneidae	"Testudinaria papposa n. gen. n. sp." = Testudinaroides papposa¹ "Testudinaria papposa n. gen. n. sp." = Testudinaroides papposa¹ Zhang et al. 1994 orb-weaving spider K0099
Insecta
Diptera - Tipulidae	Tipula corollata n. sp.¹ Tipula corollata n. sp.¹ Zhang et al. 1994 crane fly K0381
Diptera - Bibionidae	Plecia capitata n. sp.¹ Plecia capitata n. sp.¹ Zhang et al. 1994 love bug K0266
Diptera - Asilidae	Promachus paucinervis n. sp.¹ Promachus paucinervis n. sp.¹ Zhang et al. 1994 robber fly K0292
Odonata - Aeshnidae	"Mediaeschna lucida" = Epiaeschna lucida¹ "Mediaeschna lucida" = Epiaeschna lucida¹ Zhang 1989 darner dragonfly K0001
Hemiptera - Lygaeidae	Lygaeus daulus n. sp.¹ Lygaeus daulus n. sp.¹ Zhang et al. 1994 milkweed bug K0207
Hemiptera - Coreidae	Plinachtus fossilis n. sp.², Cloresmus ambimodestus n. sp.² Plinachtus fossilis n. sp.² Zhang and Zhang 1990 leaf-footed bug K0112 Cloresmus ambimodestus n. sp.² Zhang and Zhang 1990 leaf-footed bug K0083
Hemiptera - Pentatomidae	Carbula aff. crassiventris² Carbula aff. crassiventris² Dallas 1849 stink bug K0008
Hemiptera - Reduviidae	Reduvius piceus n. sp.¹, Reduvius diatomus n. sp.², Reduvius shandongianus n. sp.² Reduvius piceus n. sp.¹ Zhang et al. 1994 assassin bug K0331 Reduvius diatomus n. sp.² Zhang and Zhang 1990 assassin bug K0136 Reduvius shandongianus n. sp.² Zhang and Zhang 1990 assassin bug K0152, 0129
Hemiptera - Belostomatidae	Sphaerodema microcephalum n. sp.¹ Sphaerodema microcephalum n. sp.¹ Zhang et al. 1994 giant water bug K0359
Hemiptera - Nepidae	Ranatra dormientis¹ Ranatra dormientis¹ Zhang et al. 1994 true bug K0378
Hemiptera - Cicadidae	"Meimuna miocenica n. sp." = Cryptotympana miocenica², "Meimuna incasa n. sp." = Cryptotympana miocenica¹ "Meimuna miocenica n. sp." = Cryptotympana miocenica² Zhang and Zhang 1990 cicada K0003 "Meimuna incasa n. sp." = Cryptotympana miocenica¹ Zhang and Zhang 1990 cicada K0431
Blattodea - Blattidae	Periplaneta sphodra n. sp.¹ Periplaneta sphodra n. sp.¹ Zhang et al. 1994 cockroach K0387
Hymenoptera - Ichneumonidae	Miolyta volcania n. gen. n. sp. Miolyta volcania n. gen. n. sp. Zhang 1989 ichneumon wasp K0147
Hymenoptera - Vespidae	Vespa ciliata n. sp.¹, Provespa nobilis n. sp.¹, Harmorhynchium reflexum n. gen. n. sp.¹ Vespa ciliata n. sp.¹ Zhang et al. 1994 wasp K0371 Provespa nobilis n. sp.¹ Zhang et al. 1994 wasp K0380 Harmorhynchium reflexum n. gen. n. sp.¹ Zhang et al. 1994 potter wasp K0367
Hymenoptera - Tiphiidae	Tiphia dimidiata n. sp.¹ Tiphia dimidiata n. sp.¹ Zhang et al. 1994 wasp K0372
Hymenoptera - Formicidae	Formica flavifemoralis n. sp.¹, Camponotus pictus n. sp.¹, Alloiomma differentialis n. sp.¹, Dolichoderus luridivenosus n. sp.¹, Liometopum lubricum n. sp.¹ Formica flavifemoralis n. sp.¹ Zhang et al. 1994 ant K0267 Camponotus pictus n. sp.¹ Zhang et al. 1994 ant K0174 Alloiomma differentialis n. sp.¹ Zhang et al. 1994 ant K0288 Dolichoderus luridivenosus n. sp.¹ Zhang et al. 1994 ant K0231 Liometopum lubricum n. sp.¹ Zhang et al. 1994 ant K0295
Hymenoptera - Megachilidae	Anthidium basalticum n. sp. Anthidium basalticum n. sp. Zhang 1989 mason bee K0138
Hymenoptera - Apidae	Bombus anacolus n. sp.¹, Xylocopa obata n. sp.¹ Bombus anacolus n. sp.¹ Zhang et al. 1994 bumble bee K0347 Xylocopa obata n. sp.¹ Zhang et al. 1994 large carpenter bee K0390
Hymenoptera - Tenthredinidae	Driocampus shanwanganus n. gen. n. sp.³, Tenthredo miocenica n. sp.³, Tenthredo petrae n. sp.³ Driocampus shanwanganus n. gen. n. sp.³ Zhang and Zhang 1990 wood wasp K0057 Tenthredo miocenica n. sp.³ Zhang and Zhang 1990 wood wasp K0130 Tenthredo petrae n. sp.³ Zhang and Zhang 1990 wood wasp K0115
Hymenoptera - Cimbicidae	"Sinocimbex silacea n. sp." = Sinocimbex silaceus¹, Abia cf. lonicerae¹, "Clavellaria shandongensis" = Abia shandongensis¹ "Sinocimbex silacea n. sp." = Sinocimbex silaceus¹ Zhang et al. 1994 cimbicid sawfly K0271 Abia cf. lonicerae¹ Linnaeus 1758 cimbicid sawfly K0274 "Clavellaria shandongensis" = Abia shandongensis¹ Hong and Wang 1985 cimbicid sawfly K0644
Coleoptera - Carabidae	Rhopalochlaenius phalanthus n. gen. n. sp.¹ Rhopalochlaenius phalanthus n. gen. n. sp.¹ Zhang et al. 1994 ground beetle K0200
Coleoptera - Scarabaeidae	Petulantis yimengensis n. gen. n. sp.¹, "Dolichopoda extumida n. gen. n. sp." = Zhangsunia extumida¹, Anomala orcina n. sp.¹, Anomala endoxa n. sp.¹, Anomala brachytarsia n. sp.¹, Anomala ursa n. sp.¹, Anomala furva¹, Holotrichia cressona n. sp.¹, Adoretus rhinus n. sp.¹ Petulantis yimengensis n. gen. n. sp.¹ Zhang et al. 1994 scarab beetle K0286 "Dolichopoda extumida n. gen. n. sp." = Zhangsunia extumida¹ Zhang et al. 1994 scarab beetle K0299 Anomala orcina n. sp.¹ Zhang et al. 1994 scarab beetle K0187 Anomala endoxa n. sp.¹ Zhang et al. 1994 scarab beetle K0305 Anomala brachytarsia n. sp.¹ Zhang et al. 1994 scarab beetle K0220 Anomala ursa n. sp.¹ Zhang et al. 1994 scarab beetle K0377 Anomala furva¹ Hong 1984 scarab beetle K0049 Holotrichia cressona n. sp.¹ Zhang et al. 1994 scarab beetle K0056/0085 Adoretus rhinus n. sp.¹ Zhang et al. 1994 scarab beetle K0168
Coleoptera - Cleridae	Titanoclerus shanwangianus n. gen. n. sp. Titanoclerus shanwangianus n. gen. n. sp. Zhang 1989 checkered beetle K0126
Coleoptera - Cerambycidae	Cerambycidae indet.¹, Hornia penisularis n. gen. n. sp.¹, Podozoodes campalus n. gen. n. sp.¹, Stromatium antiquum n. sp.¹, Zoodes torridus n. sp.¹, Glischra aestivalis n. gen. n. sp.¹, Perimesosa dissita n. sp.¹, Paraphilus rejectus n. sp.¹, Mesosa longiuscula n. sp.¹, Mesocacia strumulosa n. sp.¹, Eutetrapha terenia n. sp.¹ Cerambycidae indet.¹ Latreille 1802 longhorned beetle K0277 Hornia penisularis n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0081 Podozoodes campalus n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0293 Stromatium antiquum n. sp.¹ Zhang et al. 1994 longhorned beetle K0004 Zoodes torridus n. sp.¹ Zhang et al. 1994 longhorned beetle K0366 Glischra aestivalis n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0224, 0259 Perimesosa dissita n. sp.¹ Zhang et al. 1994 longhorned beetle K0249 Paraphilus rejectus n. sp.¹ Zhang et al. 1994 longhorned beetle K0355 Mesosa longiuscula n. sp.¹ Zhang et al. 1994 longhorned beetle K0206 Mesocacia strumulosa n. sp.¹ Zhang et al. 1994 longhorned beetle K0229 Eutetrapha terenia n. sp.¹ Zhang et al. 1994 longhorned beetle K0297
Coleoptera - Buprestidae	Eolampra gorgia n. sp.¹, Illolampra ampulla n. gen. n. sp.¹, Illolampra phlegma n. sp.¹ Eolampra gorgia n. sp.¹ Zhang et al. 1994 jewel beetle K0215 Illolampra ampulla n. gen. n. sp.¹ Zhang et al. 1994 jewel beetle K0051 Illolampra phlegma n. sp.¹ Zhang et al. 1994 jewel beetle K0361
Coleoptera - Dascillidae	Lyprodascillus peritelus n. sp.¹, Dascillus musculus n. sp.¹ Lyprodascillus peritelus n. sp.¹ Zhang et al. 1994 soft bodied plant beetle K0208 Dascillus musculus n. sp.¹ Zhang et al. 1994 soft bodied plant beetle K0343
Coleoptera - Elateridae	"Corymbites euprepes n. sp." = Ctenicera euprepes¹ "Corymbites euprepes n. sp." = Ctenicera euprepes¹ Zhang et al. 1994 click beetle K0253
Coleoptera - Cantharidae	"Curticantharis trapezialis n. sp." = Themus (Themus) trapezialis¹ "Curticantharis trapezialis n. sp." = Themus (Themus) trapezialis¹ Zhang et al. 1994 soldier beetle K0349

¹Zhang et al. 1994; ²Zhang and Zhang 1990; ³Zhang and Zhang 1990