Bushy Tailed Blowout (Cretaceous of the United States)

Also known as UCMP V-5711 (BTB locality)

Where: Niobrara County, Wyoming (43.2° N, 104.6° W: paleocoordinates 49.6° N, 79.4° W)

• coordinate based on unpublished field data

• small collection-level geographic resolution

When: Lance Formation, Lancian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithified, sandy shale

• "a sandy shale"

Size class: mesofossils

Collected by W. A. Clemens in 1985; reposited in the UCMP

Primary reference: W. A. Clemens. 1964. Fossil mammals of the type Lance Formation, Wyoming. Part I. Introduction and Marsupialia. University of California Publications in Geological Sciences 48:1-105 [J. Alroy/J. Alroy/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14519: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano, Roger Benson, Mark Bell, Grace Varnham and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Bivalvia
 Unionida - Unionidae
Unionidae indet.9 Rafinesque 1820 freshwater mussel
 Cardiida - Corbiculidae
? Corbiculidae indet.9 Gray 1847 clam
Gastropoda
 Heterostropha - Physidae
cf. Physa sp.9 Draparnaud 1801 snail
 Architaenioglossa - Viviparidae
cf. Lioplacodes sp.9 Meek and Hayden 1864 snail
cf. Viviparus sp.9 Montfort 1810 snail
 Sorbeoconcha - Hydrobiidae
Reesidella sp.9 Yen 1952 snail
Chondrichthyes
 Hybodontiformes - Lonchidiidae
Lonchidion selachos9 Estes 1964 elasmobranch
UCMP 53905-53906
 Rajiformes - Sclerorhynchidae
Ischyrhiza avonicola9 Estes 1964 ray
UCMP unnumbered
 Rajiformes - Rhinobatidae
Myledaphus bipartitus9 Cope 1876 guitarfish
Reptilia
 Testudinata -
Compsemys victa9 Leidy 1856 turtle
including UCMP 173375 listed by Holroyd & Hutchison (2002)
 Testudinata - Baenidae
Eubaena cephalica9 Hay 1904 turtle
including UCMP 73030 listed by Holroyd & Hutchison (2002)
cf. Baena sp.9 Leidy 1870 turtle
 Testudines - Macrobaenidae
cf. Macrobaenidae sp.18 turtle
Voucher: UCMP 173415
 Testudines - Emydidae
Emydinae indet.9 Rafinesque 1815 turtle
"near Pseudemys?"
 Testudines - Nanhsiungchelyidae
Basilemys sp.9 Hay 1902 turtle
 Testudines - Trionychidae
Trionychidae indet.9, "Aspideretes beecheri" = Trionyx beecheri9
Trionychidae indet.9 Gray 1825 softshell turtle
costal, UCMP unnumbered; plus UCMP 173409 listed by Holroyd & Hutchison (2002)
"Aspideretes beecheri" = Trionyx beecheri9 Hay 1904 softshell turtle
 Testudines - Chelydridae
Chelydridae sp.18 Swainson 1839 turtle
Vouchers: UCMP 125773–775
 Testudines - Kinosternoidae
Kinosternoidae sp.18 Holroyd and Hutchison 2002 turtle
Voucher: UCMP 173416
 Testudines -
"Emarginochelys sp." = Emarginachelys18
"Emarginochelys sp." = Emarginachelys18 Whetstone 1978 turtle
Vouchers: UCMP 125768–772
 Testudines - Pantrionychidae
"Plastomeninae sp." = Plastomenidae18
"Plastomeninae sp." = Plastomenidae18 Hay 1902 turtle
Voucher: UCMP 173418
 Testudines - Adocidae
Adocus sp.18 Cope 1868 turtle
Voucher: UCMP 173412
 Choristodera -
Champsosaurus sp.8 Cope 1876 choristodere
 Pterosauria -
? Pterosauria indet.9 Kaup 1834 pterosaur
 Theropoda -
Theropoda indet.4 theropod
UCM 39503
? Theropoda indet.9 theropod
"straight-sided teeth"
 Theropoda - Dromaeosauridae
Dromaeosauridae indet.4 Colbert and Russell 1969 maniraptoran
UCM 39502; AMNH uncat. (n. gen. n. sp.)
Dromaeosaurinae sp.21 Matthew and Brown 1922 maniraptoran
Richardoestesia sp.21 Currie et al. 1990 maniraptoran
 Theropoda - Troodontidae
"Saurornithoides inequalis" = Stenonychosaurus inequalis4, Pectinodon bakkeri n. gen. n. sp.4
"Saurornithoides inequalis" = Stenonychosaurus inequalis4 Sternberg 1932 maniraptoran
UCM 41666, 43218
Pectinodon bakkeri n. gen. n. sp.4 Carpenter 1982 maniraptoran
UCM 38445-38446; AMNH 27117-8, 21874
Aves
 Charadriiformes -
Charadriiformes "indet. sp. 1"20 Huxley 1867 shorebird
AMNH 13011
 Charadriiformes - Cimolopterygidae
Cimolopteryx maxima n. sp.2, "Cimolopteryx petra n. sp." = Lamarqueavis petra20
Cimolopteryx maxima n. sp.2 Brodkorb 1963 shorebird
UCMP 53973, 53957
"Cimolopteryx petra n. sp." = Lamarqueavis petra20 Hope 2002 shorebird
AMNH 21911
 Charadriiformes -
Graculavus augustus n. sp.19 Hope 1999 shorebird
AMNH 25223
 Neornithes -
Neornithes "indet. sp. 6"20 bird
AMNH 25221
 Suliformes - Phalacrocoracidae
Phalacrocoracidae "indet. sp. 2"20 Reichenbach 1850 cormorant
AMNH 25272
 Neornithes -
Lonchodytes pterygius8 Brodkorb 1963 bird
 Anseriformes - Presbyornithidae
Presbyornithidae "indet. sp. 4"20 Wetmore 1926 waterfowl
AMNH 22603
Presbyornithidae "indet. sp. 3"20 Wetmore 1926 waterfowl
AMNH 21929
Presbyornithidae "indet. sp. 2"20 Wetmore 1926 waterfowl
AMNH 22602
Reptilia
 Theropoda -
Paronychodon lacustris9 Cope 1876 maniraptoran
 Theropoda - Tyrannosauridae
Aublysodon mirandus4 Leidy 1868 tyrannosaurid
 Ornithischia - Hadrosauridae
Hadrosauridae indet.4 Cope 1869 hadrosaurid
 Ornithischia -
Thescelosaurus sp.4 Gilmore 1913 ornithopod
 Ornithischia - Ceratopsidae
Ceratopsidae indet.4 Marsh 1888 ceratopsid
 Crocodylia - Alligatoridae
Brachychampsa montana9 Gilmore 1911 crocodilian
 Crocodylia -
"Leidyosuchus sternbergi" = Borealosuchus sternbergii9
"Leidyosuchus sternbergi" = Borealosuchus sternbergii9 Gilmore 1910 crocodilian
 Squamata -
Alethinophidia "Lance snake"22 snake
Fig. 1J: Lance alethinophidian vertebra
Coniophis precedens23 Marsh 1892 snake
AMNH  22846,  22850,  22851 (maxillae); AMNH 14598, 15621, 15622, 22840‐22845, 22847‐22849, 22940, 25220,  25261, 26817‐26819, 26821‐26824, 26827‐26834, 26999, 27000 (vertebrae) and UCMP  49859, 53930‐53934, 53936‐53944, 73044 (vertebrae)
 Squamata - Xenosauridae
Exostinus lancensis9 Gilmore 1928 squamates
UCMP 49847-49850, 49864, 56093; plus specimens reported by Gao & Fox (1996), UALVP 29847 (parietal), 29849-29850 (maxilla), 29838 (dentary)
 Squamata -
Paraderma bogerti9 Estes 1964 squamates
UCMP 54199, 54213
Parasaniwa wyomingensis9 Gilmore 1928 squamates
UCMP 49792, 49796, 49799, 54200; AMNH 8103; plus Gao & Fox (1996) specimens: UALVP 33351-33353 (maxilla), 33354 (frontal), 1612, 1616, 33355, 33356 (dentaries)
Litakis gilmorei9 Estes 1964 squamates
UCMP 49798
Varanoidea indet.13 Gray 1827 squamates
caudal; UCMP unnumbered
Colpodontosaurus cracens13 Estes 1964 squamates
UALVP 29781-29784 (dentaries)
 Squamata - Anguidae
"Peltosaurus piger" = Odaxosaurus piger9
"Peltosaurus piger" = Odaxosaurus piger9 Gilmore 1928 squamates
UCMP 49761-49790, 49821, 54204, 54208
 Squamata - Chamopsiidae
Leptochamops denticulatus9 Gilmore 1928 squamates
UCMP 49825-49826, 49830, 49832
Chamops segnis9 Marsh 1892 squamates
UCMP 49811-49814, 54206; and specimen reported by Gao & Fox 1996; UALVP 29727, 29729, 29813 (maxillae), and 29814 (dentary)
Meniscognathus altmani9 Estes 1964 squamates
UCMP 49837, 49841, 54266; and specimens reported by Gao & Fox 1996; UALVP 29774, 29777, 29834 (dentaries), and 29832 (maxilla)
Haptosphenus placodon9 Estes 1964 squamates
UCMP 49844-49846, 54201, 54209
 Squamata -
Lonchisaurus trichurus n. gen. n. sp.22, "? Sauriscus cooki" = Estescincosaurus cooki9, Pariguana lancensis n. gen. n. sp.22
Lonchisaurus trichurus n. gen. n. sp.22 Longrich et al. 2012 squamates
Holotype AMNH 15446, dentary
"? Sauriscus cooki" = Estescincosaurus cooki9 Estes 1964 squamates
"scincid-like parietals", UCMP
Pariguana lancensis n. gen. n. sp.22 Longrich et al. 2012 squamates
Holotype AMNH 22208, dentary
Mammalia
 Multituberculata - Cimolodontidae
Cimolodon nitidus Marsh 1889 multituberculate
 Multituberculata - Neoplagiaulacidae
Mesodma formosa Marsh 1889 multituberculate
confirmed by Novacek and Clemens 1977
Mesodma thompsoni Clemens 1964 multituberculate
confirmed by Novacek and Clemens 1977
Mesodma hensleighi24 Lillegraven 1969 multituberculate
 Multituberculata - Cimolomyidae
Cimolomys gracilis Marsh 1889 multituberculate
Meniscoessus robustus Marsh 1889 multituberculate
 Multituberculata -
Cimexomys minor7 Sloan and Van Valen 1965 multituberculate
one "Mesodma" M1 (perhaps M. "formosa") of Clemens 1964a
 Tribosphenida - Herpetotheriidae
"Nortedelphys intermedius" = Nortedelphys jasoni5
"Nortedelphys intermedius" = Nortedelphys jasoni5 Storer 1991 marsupial
 Tribosphenida - Stagodontidae
Didelphodon vorax6 Marsh 1889 metatherian
 Polydolopimorphia - Glasbiidae
Glasbius intricatus n. sp.6 Clemens 1966 metatherian
 Tribosphenida - Pediomyidae
Pediomys elegans6, "Pediomys florencae" = Protolambda florencae6, "Pediomys hatcheri" = Protolambda hatcheri6, "Pediomys krejcii" = Leptalestes krejcii6, "Pediomys cooki n. sp." = Leptalestes cooki6
Pediomys elegans6 Marsh 1889 metatherian
"Pediomys florencae" = Protolambda florencae6 Clemens 1966 metatherian
"Pediomys hatcheri" = Protolambda hatcheri6 Osborn 1898 metatherian
"Pediomys krejcii" = Leptalestes krejcii6 Clemens 1966 metatherian
this is NOT the type locality, see Clemens 1973
"Pediomys cooki n. sp." = Leptalestes cooki6 Clemens 1966 metatherian
 Tribosphenida - Alphadontidae
Alphadon marshi6 Simpson 1927 metatherian
includes A. marshi sensu stricto according to Clemens 1973; A. "wilsoni" present according to Lillegraven 1969
Turgidodon rhaister6 Clemens 1966 metatherian
 Tribosphenida -
Protalphadon lulli6 Clemens 1966 metatherian
 Creodonta -
"Cimolestes magnus" = Altacreodus magnus7
"Cimolestes magnus" = Altacreodus magnus7 Clemens and Russell 1965 creodont
 Cimolesta - Cimolestidae
Cimolestes stirtoni n. sp.7 Clemens 1973 eutherian
UCMP 48591 - type (left upper molar, probably M2)
Cimolestes incisus7 Marsh 1889 eutherian
Batodon tenuis7 Marsh 1892 eutherian
 Leptictida - Gypsonictopidae
Gypsonictops hypoconus7 Simpson 1927 eutherian
Amphibia
 Salientia -
Paradiscoglossus americanus n. gen. n. sp.11
Paradiscoglossus americanus n. gen. n. sp.11 Estes and Sanchíz 1982 frog
UCMP 125827-125829
 Salientia - Pelobatidae
cf. Eopelobates sp.15 Parker 1929 spadefoot toad
 Salientia - Discoglossidae
cf. Discoglossidae indet.11 Günther 1859 painted frog
UCMP 55706
 Salientia - Palaeobatrachidae
Palaeobatrachus occidentalis11 Estes and Sanchíz 1982 frog
UCMP 55705
 Salientia -
Neobatrachia indet.9 frog
ilium
Theatonius lancensis n. gen. n. sp.12 Fox 1976 frog
UALVP 12073-80, 40162
 Caudata - Urodela
Piceoerpeton naylori16 Gardner 2012 salamander
atlas
Lisserpeton bairdi10 Estes 1965 salamander
Scapherpeton tectum9 Cope 1876 salamander
500+ specimens, unnumbered UCMP
Habrosaurus dilatus9 Gilmore 1928 siren
UCMP unnumbered
 Proteida - Batrachosauroididae
Batrachosauroididae indet.17, Opisthotriton kayi9, "Prodesmodon copei" = Prodesmodon copei9, "Cuttysarkus mcnallyi" = Prodesmodon copei9
Batrachosauroididae indet.17 Auffenberg 1958 salamander
AMNH FARB 22965 (incomplete left dentary) - probable new genus and species,
Opisthotriton kayi9 Auffenberg 1961 salamander
UCMP unnumbered
"Prodesmodon copei" = Prodesmodon copei9 Estes 1964 salamander
UCMP unnumbered
"Cuttysarkus mcnallyi" = Prodesmodon copei9 Estes 1964 salamander
UCMP 54258
 Temnospondyli - Albanerpetontidae
Albanerpetontidae indet.14 Fox and Naylor 1982 tetrapod
AMNH 22952-22954,
Albanerpeton nexuosus14 Estes 1981 tetrapod
AMNH 15259, 22950, 22952, 22955, 22959, 27177
Actinopteri
 Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris9 Lambe 1902
UCP 54000, multiple fragments
 Teleostei -
Teleostei "indet. U-4"1
UALVP 58847, abdominal centra
Osteoglossomorpha indet.1 Greenwood et al. 1966
= Coriops/Lopadichthys; UALVP 58845, centra of large size; UALVP 58846, centra of small size
 Hiodontiformes - Hiodontidae
 Osteoglossiformes -
Wilsonichthys aridinsulensis1 Murray et al. 2016
UALVP 56054, centra type BvE
aff. Lopadichthys sp.1 Murray et al. 2018
UALVP 56055, dentary
 Esociformes - Esocidae
Estesesox sp.1 Wilson et al. 1992
UALVP 58852, dentary; UALVP 58854, palatine; UALVP 58844, centrum type NvC
 Acanthomorphata -
"Acanthomorpha "indet. HC-1"" = Acanthomorphata1, "Acanthomorpha "indet. HC-2"" = Acanthomorphata1, "Acanthomorpha "indet. HC-3"" = Acanthomorphata1, "Acanthomorpha "indet. HC-4"" = Acanthomorphata1, "Acanthomorpha "indet. HC-6"" = Acanthomorphata1, "Acanthomorpha "indet. HC-5"" = Acanthomorphata1
"Acanthomorpha "indet. HC-1"" = Acanthomorphata1 Rosen 1973
UALVP 58830, three first abdominal centra; UALVP 58876, 18 abdominal centra
"Acanthomorpha "indet. HC-2"" = Acanthomorphata1 Rosen 1973
UALVP 58832, six first abdominal centra; UALVP 58831, 14 abdominal centra
"Acanthomorpha "indet. HC-3"" = Acanthomorphata1 Rosen 1973
UALVP 58833, seven first abdominal centra
"Acanthomorpha "indet. HC-4"" = Acanthomorphata1 Rosen 1973
UALVP 58836, one first abdominal centrum; UALVP 58834, seven abdominal centra
"Acanthomorpha "indet. HC-6"" = Acanthomorphata1 Rosen 1973
UALVP 58847, one abdominal centrum
"Acanthomorpha "indet. HC-5"" = Acanthomorphata1 Rosen 1973
UALVP 58829, first abdominal centrum
 Teleostei -
Percopsiformes indet.1 Berg 1940
UALVP 59912, dentary
 Acanthomorphata - Sciaenidae
Platacodon nanus9 Marsh 1889
dentaries; UALVP 58851, pharyngeal tooth plate.
 Ostariophysi -
"Otophysi "indet. U-3/BvD"" = Otophysa1
"Otophysi "indet. U-3/BvD"" = Otophysa1
UALVP 58839, seven anterior Weberian centra (centrum type U3); UALVP 58840, two possible fourth Weberian centra; UALVP 58838, two abdominal centra (centrum type BvD)
 Teleostei -
Gonorhynchiformes "indet. H"1
UALVP 58842, eight abdominal centra (centrum type HvA)
Euteleostei "indet. RvB"1 Rosen 1973
UALVP 58848, one abdominal centrum
 Albuliformes - Albulidae
Albulidae indet.9 Bleeker 1859 bonefish
fragments
Coriops sp.1 Estes 1969 bonefish
UALVP 58853, basibranchial element of small size
 Teleostei -
Elopiformes indet.1 Sauvage 1875
UALVP 58837, one complete and one partial centrum
 Elopiformes - Phyllodontidae
Phyllodontidae indet.1 Sauvage 1875
UALVP58837
 Elopiformes - Elopidae
Elopidae indet.9 Bonaparte 1846
UCMP 55816-55817
 Elopiformes -
Ostariostoma sp.1 Schaeffer 1949
UALVP 56053, four dentaries represented by their anterior end and one complete dentary
 Amiiformes - Amiidae
cf. Protamia sp.9 Leidy 1873 bowfin
UCMP 56276
"Kindleia fragosa" = Cyclurus fragosus9 Jordan 1927 bowfin
UCMP 54045-54048, 54062, 54073-54074, 54082, 54088, 54092-54093, 54104-54105, 54178-54180, 54188, 54196-54198, 54260, 54262, 56019; + 3 + many unnumbered specimens
Melvius thomasi3 Bryant 1988 bowfin
 Lepisosteiformes - Lepisosteidae
Lepisosteus occidentalis9 Leidy 1856 gar
Osteichthyes
 Acipenseriformes - Acipenseridae
Acipenser eruciferus9 Cope 1876 sturgeon
UCMP unnumbered