Swain Quarry (Paleocene of the United States)

Also known as CM loc 217

Where: Carbon County, Wyoming (41.3° N, 107.7° W: paleocoordinates 47.7° N, 86.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Fort Union Formation, Torrejonian (63.8 - 60.9 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size classes: macrofossils, mesofossils

Collection methods: surface (in situ),

Primary reference: R. M. West. 1970. Tetraclaenodon puercensis (Mammalia: Phenacodontidae), Goler Formation, Paleocene of California, and distribution of the genus. Journal of Paleontology 44(5):851-857 [J. Alroy/J. Alroy/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14886: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Roger Benson, Terri Cleary and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• there are seven other localities in the same section, but none of the material is described

•Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977

Mammalia
 Multituberculata - Ptilodontidae
Ptilodus mediaevus9 Cope 1881 multituberculate
Baiotomeus douglassi8 Simpson 1935 multituberculate
one specimen of "Ptilodus mediaevus"
 Multituberculata - Neoplagiaulacidae
"Parectypodus clemensi" = Krauseia clemensi12, Neoplagiaulax nelsoni13, Xanclomys mcgrewi n. sp.9, "Ectypodus sylviae n. sp." = Parectypodus sylviae9, Xyronomys swainae n. sp.9, Ectypodus szalayi12
"Parectypodus clemensi" = Krauseia clemensi12 Sloan 1981 multituberculate
may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan
Neoplagiaulax nelsoni13 Sloan 1987 multituberculate
N. "cf. hunteri"
Xanclomys mcgrewi n. sp.9 Rigby 1980 multituberculate
"Ectypodus sylviae n. sp." = Parectypodus sylviae9 Rigby 1980 multituberculate
Xyronomys swainae n. sp.9 Rigby 1980 multituberculate
Ectypodus szalayi12 Sloan 1981 multituberculate
 Cimolesta - Apatemyidae
Jepsenella praepropera9 Simpson 1940 eutherian
 Cimolesta - Cimolestidae
Paleotomus milleri n. sp.9 Rigby 1980 eutherian
Gelastops joni n. sp.9 Rigby 1980 eutherian
 Taeniodonta - Stylinodontidae
Psittacotherium multifragum9 Cope 1882 taeniodont
 Proteutheria - Palaeoryctidae
Palaeoryctes puercensis9 Matthew 1913 eutherian
 Insectivora -
Insectivora indet.1 Bowdich 1821 insectivoran
 Condylarthra - Hyopsodontidae
Litaletes sp.10 Simpson 1935 condylarth
said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus
Promioclaenus acolytus9 Cope 1882 condylarth
 Placentalia -
 Macroscelidea -
"Mckennatherium martinezi n. sp." = Adunator martinezi9, "Mckennatherium fredericki n. sp." = Adunator fredericki9
"Mckennatherium martinezi n. sp." = Adunator martinezi9 Rigby 1980 elephant shrew
"Mckennatherium fredericki n. sp." = Adunator fredericki9 Rigby 1980 elephant shrew
 Macroscelidea - Apheliscidae
Litomylus dissentaneus9 Simpson 1935 elephant shrew
Haplaletes disceptatrix9 Simpson 1935 elephant shrew
 Tribosphenida - Mixodectidae
Mixodectes sp.9 Cope 1883 placental
Mixodectes malaris9 Cope 1884 placental
confirmed by Gunnell 1989
 Primates - Palaechthonidae
Palenochtha minor6, Palenochtha weissae n. sp.9, "Palaechthon problematicus" = Plesiolestes problematicus9
Palenochtha minor6 Gidley 1923 primate
Palenochtha weissae n. sp.9 Rigby 1980 primate
"Palaechthon problematicus" = Plesiolestes problematicus9 Jepsen 1930 primate
 Primates - Paromomyidae
Paromomys depressidens9 Gidley 1923 primate
confirmed by Gunnell 1989
Paromomys maturus9 Gidley 1923 primate
confirmed by Gunnell 1989
 Primates - Picrodontidae
Picrodus silberlingi9 Douglass 1908 primate
 Tribosphenida -
"Prodiacodon cf. puercensis" = Diacodon puercensis9
"Prodiacodon cf. puercensis" = Diacodon puercensis9 Matthew 1918 placental
 Soricomorpha - Nyctitheriidae
Leptacodon cf. tener9 Matthew and Granger 1921 placental
 Tribosphenida - Pantolestoidae
Bessoecetor septentrionalis11 Russell 1929 placental
Pantolestinae indet.9 Simpson 1937 placental
 Acreodi - Mesonychidae
Dissacus navajovius9 Cope 1881 condylarth
 Acreodi - Oxyclaenidae
Prothryptacodon hilli n. sp.9 Rigby 1980 placental
Prothryptacodon cf. furens9 Simpson 1935 placental
 Acreodi - Triisodontidae
Goniacodon levisanus9 Cope 1883 placental
 Tribosphenida - Phenacodontidae
Tetraclaenodon puercensis9 Cope 1881 condylarth
 Procreodi - Arctocyonidae
Chriacus pelvidens9, "Mimotricentes subtrigonus" = Tricentes subtrigonus9, Arctocyon ferox9, "Protogonodon criswelli n. sp." = Loxolophus criswelli9
Chriacus pelvidens9 Cope 1881 condylarth
"Mimotricentes subtrigonus" = Tricentes subtrigonus9 Cope 1881 condylarth
Arctocyon ferox9 Cope 1883 condylarth
"Protogonodon criswelli n. sp." = Loxolophus criswelli9 Rigby 1980 condylarth
 Ferae - Viverravidae
"Bryanictis vanvaleni" = Intyrictis vanvaleni9, "Protictis haydenianus" = Protictis haydenianus9, "Prolimnocyon macfaddeni n. sp." = Protictis haydenianus9, Simpsonictis jaynanneae n. sp.9
"Bryanictis vanvaleni" = Intyrictis vanvaleni9 MacIntyre 1966 placental
"Protictis haydenianus" = Protictis haydenianus9 Cope 1882 placental
"Prolimnocyon macfaddeni n. sp." = Protictis haydenianus9 Cope 1882 placental
macfaddeni
Simpsonictis jaynanneae n. sp.9 Rigby 1980 placental
 Tribosphenida - Periptychidae
Periptychus carinidens9 Cope 1881 condylarth
Anisonchus willeyi n. sp.9 Rigby 1980 condylarth
Anisonchus sectorius9 Cope 1881 condylarth
Haploconus sp.9 Cope 1882 condylarth
 Leptictida - Leptictidae
"Diacodon concordiarcensis" = Prodiacodon concordiarcensis9, Myrmecoboides montanensis9, "Acmeodon hyoni n. sp." = Acmeodon secans9
"Diacodon concordiarcensis" = Prodiacodon concordiarcensis9 Simpson 1935 eutherian
Myrmecoboides montanensis9 Gidley 1915 eutherian
"Acmeodon hyoni n. sp." = Acmeodon secans9 Matthew and Granger 1921 eutherian
 Tribosphenida - Herpetotheriidae
Swaindelphys cifellii n. sp.7 Johanson 1996 marsupial
Reptilia
 Squamata -
Squamata indet.14 squamates
"indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
 Squamata - Iguanidae
Swainiguanoides milleri n. gen. n. sp.14
Swainiguanoides milleri n. gen. n. sp.14 Sullivan 1982 squamates
AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
 Squamata - Necrosauridae
cf. Provaranosaurus sp.14 Gilmore 1942 squamates
AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
 Squamata - Xenosauridae
Exostinus cf. lancensis14 Gilmore 1928 squamates
AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
 Squamata -
cf. Saniwa sp.14 Leidy 1870 squamates
AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra.
cf. Saniwa sp.4 Leidy 1870 squamates
AMNH 15957, medial part of left dentary with 3 tooth bases.
 Squamata - Anguidae
Anguidae indet.14 Gray 1825 squamates
AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries.
Odaxosaurinae indet.14 Sullivan 1982 squamates
AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm.
Glyptosaurinae indet.14 Marsh 1872 squamates
AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla
Odaxosaurus sp.2 Gilmore 1928 squamates
CM 87168
Odaxosaurus piger14 Gilmore 1928 squamates
AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary.
Machaerosaurus torrejonensis14 Gilmore 1928 squamates
AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
 Squamata - Xantusiidae
Palaeoxantusia fera14 Hecht 1956 night lizard
AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla.
Palaeoxantusia fera4 Hecht 1956 night lizard
scincid
 Testudinata - Baenidae
"Plesiobaena putorius" = Cedrobaena putorius3
"Plesiobaena putorius" = Cedrobaena putorius3 Gaffney 1972 turtle
AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof
Amphibia
 Caudata - Urodela
Habrosaurus sp.5 Gilmore 1928 siren
Scapherpeton tectum5 Cope 1876 salamander
 Proteida - Batrachosauroididae
Opisthotriton kayi5 Auffenberg 1961 salamander