PBDB

Swain Quarry (Paleocene of the United States)

Where: Carbon County, Wyoming (41.3° N, 107.7° W: paleocoordinates 47.7° N, 86.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Fort Union Formation, Torrejonian (63.3 - 61.7 Ma)

Environment/lithology: terrestrial

Size classes: macrofossils, mesofossils

Primary reference: J. K. Rigby, Jr. 1980. Swain Quarry of the Fort Union Formation, Middle Paleocene (Torrejonian), Carbon County, Wyoming: geologic setting and mammalian fauna. Evolutionary Monographs 3:1-178 [J. Alroy/J. Alroy/J. Marcot]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14886: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Roger Benson, Terri Cleary and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• there are seven other localities in the same section, but none of the material is described

•Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977

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Osteichthyes
Proteida - Batrachosauroididae	Opisthotriton kayi⁴ Opisthotriton kayi⁴ Auffenberg 1961 tetrapod
Caudata - Urodela	Habrosaurus sp.⁴, Scapherpeton tectum⁴ Habrosaurus sp.⁴ Gilmore 1928 siren Scapherpeton tectum⁴ Cope 1876 salamander
Reptilia
Testudines - Baenidae	"Plesiobaena putorius" = Cedrobaena putorius² "Plesiobaena putorius" = Cedrobaena putorius² Gaffney 1972 turtle AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof
Squamata -	Squamata indet.¹² Squamata indet.¹² Oppel 1811 lizard "indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
Squamata - Xenosauridae	Exostinus cf. lancensis¹² Exostinus cf. lancensis¹² Gilmore 1928 lizard AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
Squamata - Necrosauridae	cf. Provaranosaurus sp.¹² cf. Provaranosaurus sp.¹² Gilmore 1942 lizard AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
Squamata -	cf. Saniwa sp.¹², cf. Saniwa sp.³ cf. Saniwa sp.¹² Leidy 1870 lizard AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra. cf. Saniwa sp.³ Leidy 1870 lizard AMNH 15957, medial part of left dentary with 3 tooth bases.
Squamata - Anguidae	Anguidae indet.¹², Glyptosaurinae indet.¹², Odaxosaurus piger¹², Odaxosaurinae indet.¹², Machaerosaurus torrejonensis¹² Anguidae indet.¹² Gray 1825 lizard AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries. Glyptosaurinae indet.¹² Marsh 1872 lizard AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla Odaxosaurus piger¹² Gilmore 1928 lizard AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary. Odaxosaurinae indet.¹² Sullivan 1982 lizard AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm. Machaerosaurus torrejonensis¹² Gilmore 1928 lizard AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
Squamata - Iguanidae	Swainiguanoides milleri n. gen. n. sp.¹² Swainiguanoides milleri n. gen. n. sp.¹² Sullivan 1982 lizard AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
Squamata - Xantusiidae	Palaeoxantusia fera¹², Palaeoxantusia fera³ Palaeoxantusia fera¹² Hecht 1956 night lizard AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla. Palaeoxantusia fera³ Hecht 1956 night lizard scincid
Mammalia
Cimolesta - Cimolestidae	Paleotomus milleri n. sp., Acmeodon secans¹, Gelastops joni n. sp. Paleotomus milleri n. sp. Rigby 1980 eutherian Acmeodon secans¹ Matthew and Granger 1921 eutherian Gelastops joni n. sp. Rigby 1980 eutherian
Cimolesta - Palaeoryctidae	Palaeoryctes puercensis Palaeoryctes puercensis Matthew 1913 eutherian
Cimolesta - Pantolestidae	Pantolestinae indet., Bessoecetor septentrionalis⁹ Pantolestinae indet. Simpson 1937 eutherian Bessoecetor septentrionalis⁹ Russell 1929 eutherian
Leptictida - Leptictidae	Myrmecoboides montanensis, Prodiacodon cf. puercensis, Prodiacodon concordiarcensis¹ Myrmecoboides montanensis Gidley 1915 eutherian Prodiacodon cf. puercensis Matthew 1918 eutherian Prodiacodon concordiarcensis¹ Simpson 1935 eutherian
Primates - Picrodontidae	Picrodus silberlingi Picrodus silberlingi Douglass 1908 primate
Primates - Paromomyidae	Paromomys maturus, Paromomys depressidens Paromomys maturus Gidley 1923 primate confirmed by Gunnell 1989 Paromomys depressidens Gidley 1923 primate confirmed by Gunnell 1989
Primates - Palaechthonidae	Palenochtha minor⁵, Palenochtha weissae n. sp., Plesiolestes problematicus⁵ Palenochtha minor⁵ Gidley 1923 primate Palenochtha weissae n. sp. Rigby 1980 primate Plesiolestes problematicus⁵ Jepsen 1930 primate
Condylarthra - Hyopsodontidae	Litaletes sp.⁸, Promioclaenus acolytus Litaletes sp.⁸ Simpson 1935 condylarth said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus Promioclaenus acolytus Cope 1882 condylarth
Condylarthra - Chriacidae	Prothryptacodon hilli n. sp., Prothryptacodon cf. furens Prothryptacodon hilli n. sp. Rigby 1980 placental Prothryptacodon cf. furens Simpson 1935 placental
Placentalia -	Deltatheridia indet. Deltatheridia indet. placental
Tribosphenida - Periptychidae	Periptychus carinidens, Haploconus sp., Anisonchus sectorius, Anisonchus willeyi n. sp. Periptychus carinidens Cope 1881 condylarth Haploconus sp. Cope 1882 condylarth Anisonchus sectorius Cope 1881 condylarth Anisonchus willeyi n. sp. Rigby 1980 condylarth
Tribosphenida - Phenacodontidae	Tetraclaenodon puercensis Tetraclaenodon puercensis Cope 1881 condylarth
Tribosphenida - Nyctitheriidae	Leptacodon cf. tener Leptacodon cf. tener Matthew and Granger 1921 placental
Ferae - Viverravidae	Simpsonictis jaynanneae n. sp., Protictis haydenianus, Intyrictis vanvaleni¹ Simpsonictis jaynanneae n. sp. Rigby 1980 placental Protictis haydenianus Cope 1882 placental Intyrictis vanvaleni¹ MacIntyre 1966 placental
Macroscelidea - Apheliscidae	Haplaletes disceptatrix, Litomylus dissentaneus Haplaletes disceptatrix Simpson 1935 elephant shrew Litomylus dissentaneus Simpson 1935 elephant shrew
Macroscelidea -	Adunator fredericki¹, Adunator martinezi¹ Adunator fredericki¹ Rigby 1980 elephant shrew Adunator martinezi¹ Rigby 1980 elephant shrew
Taeniodonta - Stylinodontidae	Psittacotherium multifragum Psittacotherium multifragum Cope 1882 taeniodont
Tribosphenida - Arctocyonidae	Arctocyon ferox, Loxolophus criswelli¹, "Mimotricentes subtrigonus" = Tricentes subtrigonus, Chriacus pelvidens Arctocyon ferox Cope 1883 condylarth Loxolophus criswelli¹ Rigby 1980 condylarth "Mimotricentes subtrigonus" = Tricentes subtrigonus Cope 1881 condylarth Chriacus pelvidens Cope 1881 condylarth
Acreodi - Triisodontidae	Goniacodon levisanus Goniacodon levisanus Cope 1883 eutherian
Acreodi - Mesonychidae	Dissacus navajovius Dissacus navajovius Cope 1881 condylarth
Theriiformes - Herpetotheriidae	Swaindelphys cifellii n. sp.⁶ Swaindelphys cifellii n. sp.⁶ Johanson 1996 marsupial
Insectivora -	Insectivora indet.¹ Insectivora indet.¹ insectivoran
Sarcobora - Mixodectidae	Mixodectes sp., Mixodectes malaris Mixodectes sp. Cope 1883 insectivoran Mixodectes malaris Cope 1884 insectivoran confirmed by Gunnell 1989
Sarcobora - Apatemyidae	Jepsenella praepropera Jepsenella praepropera Simpson 1940 insectivoran
Multituberculata - Neoplagiaulacidae	Parectypodus clemensi¹⁰, Parectypodus sylviae¹, Ectypodus szalayi¹⁰, Xanclomys mcgrewi n. sp., Neoplagiaulax nelsoni¹¹, Xyronomys swainae n. sp. Parectypodus clemensi¹⁰ Sloan 1981 multituberculate may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan Parectypodus sylviae¹ Rigby 1980 multituberculate Ectypodus szalayi¹⁰ Sloan 1981 multituberculate Xanclomys mcgrewi n. sp. Rigby 1980 multituberculate Neoplagiaulax nelsoni¹¹ Sloan 1987 multituberculate N. "cf. hunteri" Xyronomys swainae n. sp. Rigby 1980 multituberculate
Multituberculata - Ptilodontidae	Baiotomeus douglassi⁷, Ptilodus mediaevus Baiotomeus douglassi⁷ Simpson 1935 multituberculate one specimen of "Ptilodus mediaevus" Ptilodus mediaevus Cope 1881 multituberculate

¹Alroy 2002; ²Gaffney 1972; ³Gao and Fox 1996; ⁴Gardner and DeMar 2013; ⁵Gunnell 1989; ⁶Johanson 1996; ⁷Krause 1987; ⁸Robison 1986; ⁹Scott et al. 2002; ¹⁰Sloan 1981; ¹¹Sloan 1987; ¹²Sullivan 1982