Swain Quarry (Paleocene of the United States)

Where: Carbon County, Wyoming (41.3° N, 107.7° W: paleocoordinates 47.7° N, 86.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Fort Union Formation, Torrejonian (63.3 - 61.7 Ma)

Environment/lithology: terrestrial

Size classes: macrofossils, mesofossils

Primary reference: J. K. Rigby, Jr. 1980. Swain Quarry of the Fort Union Formation, Middle Paleocene (Torrejonian), Carbon County, Wyoming: geologic setting and mammalian fauna. Evolutionary Monographs 3:1-178 [J. Alroy/J. Alroy/J. Marcot]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14886: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Roger Benson, Terri Cleary and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• there are seven other localities in the same section, but none of the material is described

•Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977

Osteichthyes
 Proteida - Batrachosauroididae
Opisthotriton kayi4 Auffenberg 1961 tetrapod
 Caudata - Urodela
Habrosaurus sp.4 Gilmore 1928 siren
Scapherpeton tectum4 Cope 1876 salamander
Reptilia
 Testudines - Baenidae
"Plesiobaena putorius" = Cedrobaena putorius2
"Plesiobaena putorius" = Cedrobaena putorius2 Gaffney 1972 turtle
AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof
 Squamata -
Squamata indet.12 Oppel 1811 lizard
"indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
 Squamata - Xenosauridae
Exostinus cf. lancensis12 Gilmore 1928 lizard
AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
 Squamata - Necrosauridae
cf. Provaranosaurus sp.12 Gilmore 1942 lizard
AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
 Squamata -
cf. Saniwa sp.12 Leidy 1870 lizard
AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra.
cf. Saniwa sp.3 Leidy 1870 lizard
AMNH 15957, medial part of left dentary with 3 tooth bases.
 Squamata - Anguidae
Anguidae indet.12 Gray 1825 lizard
AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries.
Glyptosaurinae indet.12 Marsh 1872 lizard
AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla
Odaxosaurus piger12 Gilmore 1928 lizard
AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary.
Odaxosaurinae indet.12 Sullivan 1982 lizard
AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm.
Machaerosaurus torrejonensis12 Gilmore 1928 lizard
AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
 Squamata - Iguanidae
Swainiguanoides milleri n. gen. n. sp.12
Swainiguanoides milleri n. gen. n. sp.12 Sullivan 1982 lizard
AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
 Squamata - Xantusiidae
Palaeoxantusia fera12 Hecht 1956 night lizard
AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla.
Palaeoxantusia fera3 Hecht 1956 night lizard
scincid
Mammalia
 Cimolesta - Cimolestidae
Paleotomus milleri n. sp. Rigby 1980 eutherian
Acmeodon secans1 Matthew and Granger 1921 eutherian
Gelastops joni n. sp. Rigby 1980 eutherian
 Cimolesta - Palaeoryctidae
Palaeoryctes puercensis Matthew 1913 eutherian
 Cimolesta - Pantolestidae
Pantolestinae indet. Simpson 1937 eutherian
Bessoecetor septentrionalis9 Russell 1929 eutherian
 Leptictida - Leptictidae
Myrmecoboides montanensis Gidley 1915 eutherian
Prodiacodon cf. puercensis Matthew 1918 eutherian
Prodiacodon concordiarcensis1 Simpson 1935 eutherian
 Primates - Picrodontidae
Picrodus silberlingi Douglass 1908 primate
 Primates - Paromomyidae
Paromomys maturus Gidley 1923 primate
confirmed by Gunnell 1989
Paromomys depressidens Gidley 1923 primate
confirmed by Gunnell 1989
 Primates - Palaechthonidae
Palenochtha minor5 Gidley 1923 primate
Palenochtha weissae n. sp. Rigby 1980 primate
Plesiolestes problematicus5 Jepsen 1930 primate
 Condylarthra - Hyopsodontidae
Litaletes sp.8 Simpson 1935 condylarth
said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus
Promioclaenus acolytus Cope 1882 condylarth
 Condylarthra - Chriacidae
Prothryptacodon hilli n. sp. Rigby 1980 placental
Prothryptacodon cf. furens Simpson 1935 placental
 Placentalia -
 Tribosphenida - Periptychidae
Periptychus carinidens Cope 1881 condylarth
Haploconus sp. Cope 1882 condylarth
Anisonchus sectorius Cope 1881 condylarth
Anisonchus willeyi n. sp. Rigby 1980 condylarth
 Tribosphenida - Phenacodontidae
Tetraclaenodon puercensis Cope 1881 condylarth
 Tribosphenida - Nyctitheriidae
Leptacodon cf. tener Matthew and Granger 1921 placental
 Ferae - Viverravidae
Simpsonictis jaynanneae n. sp. Rigby 1980 placental
Protictis haydenianus Cope 1882 placental
Intyrictis vanvaleni1 MacIntyre 1966 placental
 Macroscelidea - Apheliscidae
Haplaletes disceptatrix Simpson 1935 elephant shrew
Litomylus dissentaneus Simpson 1935 elephant shrew
 Macroscelidea -
Adunator fredericki1 Rigby 1980 elephant shrew
Adunator martinezi1 Rigby 1980 elephant shrew
 Taeniodonta - Stylinodontidae
Psittacotherium multifragum Cope 1882 taeniodont
 Tribosphenida - Arctocyonidae
Arctocyon ferox Cope 1883 condylarth
Loxolophus criswelli1 Rigby 1980 condylarth
"Mimotricentes subtrigonus" = Tricentes subtrigonus Cope 1881 condylarth
Chriacus pelvidens Cope 1881 condylarth
 Acreodi - Triisodontidae
Goniacodon levisanus Cope 1883 eutherian
 Acreodi - Mesonychidae
Dissacus navajovius Cope 1881 condylarth
 Theriiformes - Herpetotheriidae
Swaindelphys cifellii n. sp.6 Johanson 1996 marsupial
 Insectivora -
Insectivora indet.1 insectivoran
 Sarcobora - Mixodectidae
Mixodectes sp. Cope 1883 insectivoran
Mixodectes malaris Cope 1884 insectivoran
confirmed by Gunnell 1989
 Sarcobora - Apatemyidae
Jepsenella praepropera Simpson 1940 insectivoran
 Multituberculata - Neoplagiaulacidae
Parectypodus clemensi10 Sloan 1981 multituberculate
may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan
Parectypodus sylviae1 Rigby 1980 multituberculate
Ectypodus szalayi10 Sloan 1981 multituberculate
Xanclomys mcgrewi n. sp. Rigby 1980 multituberculate
Neoplagiaulax nelsoni11 Sloan 1987 multituberculate
N. "cf. hunteri"
Xyronomys swainae n. sp. Rigby 1980 multituberculate
 Multituberculata - Ptilodontidae
Baiotomeus douglassi7 Simpson 1935 multituberculate
one specimen of "Ptilodus mediaevus"
Ptilodus mediaevus Cope 1881 multituberculate