PBDB Collection

Princeton Quarry (Paleocene of the United States)

Also known as Big Sand Coulee; Locality No. 3; SC-187

Where: Park County, Wyoming (44.9° N, 108.9° W: paleocoordinates 50.8° N, 88.4° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Fort Union Formation, Tiffanian (61.7 - 56.8 Ma)

• low within chron C25r according to Butler et al. 1981 but in the middle of C25r according to Secord et al. 2006

•

•"Polecat Bench Fm, Silver Coulee beds" according to Gilmore 1942

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Collected by G. L. Jepsen; reposited in the UMMP

• Princeton University collection

Primary reference: K. D. Rose. 1981. The Clarkforkian Land-Mammal Age and Mammalian Faunal Composition Across the Paleocene-Eocene Boundary. University of Michigan Papers on Paleontology 26:1-197 [J. Alroy/J. Alroy/P. Mannion]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 15049: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Terri Cleary, K. Christopher Beard, Patricia Holroyd, Robin Whatley, Evangelos Vlachos and Jonathan Marcot

Creative Commons license: CC BY (attribution)

Taxonomic list

• replaces list of Jepsen and Woodburne 1969

Show authors, comments, and common names

Amphibia
Salientia -	Anura indet.² Anura indet.² Fischer von Waldheim 1813 frog PU 14662, distal end of right radioulna; 14663, proximal end of left radioulna; 14669 (part), phalanx; 13372, right maxilla.
Salientia - Discoglossidae	Discoglossidae indet.² Discoglossidae indet.² Günther 1859 painted frog PU 14670, distal end of right humerus. Undescribed genus and species.
Salientia - Rhinophrynidae	Eorhinophrynus sp.² Eorhinophrynus sp.² Hecht 1959 burrowing toad PU 14664, left humerus; 14665, distal end of left humerus; 14669 (part), sacral vertebra.
Caudata - Urodela	Amphiuma jepseni n. sp.², Scapherpeton tectum² Amphiuma jepseni n. sp.² Estes 1969 amphiuma PU 14666 (holotype), partial vertebral column with at least 14 associated but dislocated vertebrae in a single block of matrix, associated left quadrate and left dentary; 14668, partial skull and associated mandibles; 16788, group of dislocated vertebrae from sacral region of young individual mingled with vertebrae of Opisthotriton kayi Scapherpeton tectum² Cope 1876 salamander PU 20583, three broken vertebrae, an atlas, an ilium, and a rib.
Proteida - Batrachosauroididae	Opisthotriton kayi² Opisthotriton kayi² Auffenberg 1961 salamander PU 14575, articulated skeleton, including poorly-preserved skull, mandibles, atlas, at least 35 presacral vertebrae, many in natural articulation, fragments of hyobranchial skeleton, a humerus, and a single rib; PU 14643, partial skeleton of a somewhat smaller individual, including skull, mandibles, partial hyobranchial skeleton in natural articulation with skull, atlas and five associated vertebrae in natural articulation; 20584, fused articular-prearticular, otoccipitals, associated with material of Amphiuma jepseni noted below; 21781, two vertebral centra associated with a block of dislocated vertebrae, mostly young Amphiuma jepseni.
Reptilia
Crocodylia - Alligatoridae	Allognathosuchus sp.² Allognathosuchus sp.² Mook 1921 crocodilian PU 16988, anterior end of left dentary.
Squamata - Oligodontosauridae	Oligodontosaurus wyomingensis n. gen. n. sp.⁴ Oligodontosaurus wyomingensis n. gen. n. sp.⁴ Gilmore 1942 worm lizard YPM-PU 14246 (holotype), left mandible
Squamata -	"Haplodontosaurus excedens" = Harpagosaurus excedens⁴ "Haplodontosaurus excedens" = Harpagosaurus excedens⁴ Gilmore 1928 squamates
Squamata - Necrosauridae	Provaranosaurus acutus n. gen. n. sp.⁴ Provaranosaurus acutus n. gen. n. sp.⁴ Gilmore 1942 squamates PU 14243 (holotype), left maxilla; 14561, anterior portion of left dentary; 17145, questionably referred fragment of left maxilla of large specimen
Squamata -	Restes rugosus n. gen. n. sp.³ Restes rugosus n. gen. n. sp.³ Gauthier 1982 squamates PU 14559, posterior half of right maxilla (holotype); 14560, right maxilla of smaller individual; 17144, left dentary fragment; 14640, disarticulated skull, jaws, and fragments of skeleton, including maxilla, prefrontals, frontal, right jugal, left pterygoid, dentaries, right prearticular- surangular; coronoids, right angular, centrum of axis, several cervical vertebrae, fragmentary left ilium and unidentified fragments.
Squamata - Anguidae	Odaxosaurus piger⁸, "Peltosaurus jepseni n. sp." = Proxestops silberlingii⁴ Odaxosaurus piger⁸ Gilmore 1928 squamates PU 13371, left maxilla; 14562, left maxilla; 14565, right maxilla, posterior part of parietal, one dermal scute; 14574, left maxilla, anterior end of right dentary; 16790, left dentary; 17147, left dentary; 17148, left dentary fragment and coronoid; 18578, left dentary fragment and osteoscutes; 14667, right pterygoid; 16761, parietal. "Peltosaurus jepseni n. sp." = Proxestops silberlingii⁴ Gilmore 1938 squamates Recombined into Proxestops jepseni by Gauthier (1982), synonymised into Proxestops silberlingii by Sullivan (1991). PU 14565 (originally holotype); UCMP 99851, isolated osteoderms; UCMP 100163, right maxilla missing all teeth, premaxillary, posterior and nasal processes; UCMP 100164, worn fragment from posterior end of right maxilla; UCMP 115128, posterior process of left maxilla; UCMP 115130, fragment from middle of right maxilla. Some specimens here removed and reclassified as Xestops savagei by ref. 60296 in coll. 15451.
Testudines - Kinosternoidea	Cardichelyon rogerwoodi⁶ Cardichelyon rogerwoodi⁶ Hutchison 2013 mud turtle PU 17794, fragments of skull and shell.
Mammalia
Multituberculata -	Neoliotomus conventus Neoliotomus conventus Jepsen 1930 multituberculate
Multituberculata - Ptilodontidae	Prochetodon cavus n. sp. Prochetodon cavus n. sp. Jepsen 1940 multituberculate
Multituberculata - Neoplagiaulacidae	Mimetodon churchilli n. sp., Neoplagiaulax hazeni n. sp., "Parectypodus laytoni n. sp." = Ectypodus laytoni, Ectypodus powelli n. sp. Mimetodon churchilli n. sp. Jepsen 1940 multituberculate Neoplagiaulax hazeni n. sp. Jepsen 1940 multituberculate "Parectypodus laytoni n. sp." = Ectypodus laytoni Jepsen 1940 multituberculate Ectypodus powelli n. sp. Jepsen 1940 multituberculate
Multituberculata - Microcosmodontidae	Microcosmodon conus n. sp., Pentacosmodon pronus n. sp. Microcosmodon conus n. sp. Jepsen 1930 multituberculate Pentacosmodon pronus n. sp. Jepsen 1940 multituberculate
Theriamorpha - Leptictidae	? Palaeictops sp., Prodiacodon paucus n. sp.¹⁰ ? Palaeictops sp. Matthew 1899 eutherian Prodiacodon paucus n. sp.¹⁰ Secord 2008 eutherian
Cimolesta - Apatemyidae	Unuchinia sp. Unuchinia sp. Simpson 1937 eutherian
Cimolesta - Barylambdidae	Barylambdidae indet.¹⁰ Barylambdidae indet.¹⁰ Patterson 1939 pantodont
Macroscelidea -	Adunator abditus n. sp.¹⁰, "Diacocherus minutus" = Adunator minutus¹ Adunator abditus n. sp.¹⁰ Secord 2008 elephant shrew "Diacocherus minutus" = Adunator minutus¹ Jepsen 1930 elephant shrew
Macroscelidea - Apheliscidae	Phenacodaptes sabulosus n. sp., Phenacodaptes sabulosus¹⁰ Phenacodaptes sabulosus n. sp. Jepsen 1930 elephant shrew Phenacodaptes sabulosus¹⁰ Jepsen 1930 elephant shrew
Acreodi - Mesonychidae	Dissacus argenteus n. sp.⁹ Dissacus argenteus n. sp.⁹ O'Leary and Rose 1995 condylarth
Theriamorpha - Metacheiromyidae	Propalaeanodon schaffi Propalaeanodon schaffi Rose 1979 placental
Carnivora - Miacidae	Viverravinae indet. Viverravinae indet. Matthew 1909 carnivoran "Viverravine sp. B" of Rose
Theriamorpha - Viverravidae	Viverravus laytoni¹, Viverravus schaffi¹⁰, Didymictis dellensis¹⁰ Viverravus laytoni¹ Gingerich and Winkler 1985 placental Viverravus schaffi¹⁰ Gingerich and Winkler 1985 placental Didymictis dellensis¹⁰ Dorr 1952 placental
Theriamorpha - Phenacodontidae	Ectocion mediotuber n. sp.¹¹, Phenacodus grangeri¹⁰, Phenacodus vortmani¹⁰ Ectocion mediotuber n. sp.¹¹ Thewissen 1990 condylarth Phenacodus grangeri¹⁰ Simpson 1935 condylarth Phenacodus vortmani¹⁰ Cope 1880 condylarth
Procreodi - Arctocyonidae	Thryptacodon pseudarctos¹⁰, Princetonia cf. yalensis¹⁰, Arctocyon corrugatus⁷ Thryptacodon pseudarctos¹⁰ Simpson 1928 condylarth Princetonia cf. yalensis¹⁰ Gingerich 1989 condylarth Arctocyon corrugatus⁷ Cope 1883 condylarth
Theriamorpha - Nyctitheriidae	Leptacodon packi n. sp., Wyonycteris galensis n. sp.¹⁰ Leptacodon packi n. sp. Jepsen 1930 placental Wyonycteris galensis n. sp.¹⁰ Secord 2008 placental
Theriamorpha - Apternodontidae	Apternodontidae indet. Apternodontidae indet. Matthew 1910 placental "Apternodus-like" A and B of Rose
Theriamorpha - Erinaceidae	Litolestes ignotus n. sp. Litolestes ignotus n. sp. Jepsen 1930 hedgehog
Primates - Carpolestidae	Carpolestes dubius n. sp. Carpolestes dubius n. sp. Jepsen 1930 primate
Primates - Plesiadapidae	Plesiadapis fodinatus n. sp., Plesiadapis churchilli¹⁰ Plesiadapis fodinatus n. sp. Jepsen 1930 primate Plesiadapis churchilli¹⁰ Gingerich 1975 primate
Primates - Micromomyidae	Micromomys silvercouleei n. sp. Micromomys silvercouleei n. sp. Szalay 1973 primate
Primates - Paromomyidae	Dillerlemur pagei¹ Dillerlemur pagei¹ Jepsen 1930 primate
Proteutheria - Palaeoryctidae	Palaeoryctes cf. punctatus⁵ Palaeoryctes cf. punctatus⁵ Van Valen 1966 eutherian
Theriamorpha - Peradectidae	Peradectes elegans Peradectes elegans Matthew and Granger 1921 marsupial
Didelphimorphia - Didelphidae	Didelphidae indet.¹⁰ Didelphidae indet.¹⁰ opossum

¹Alroy 2002; ²Estes 1975; ³Gauthier 1982; ⁴Gilmore 1942; ⁵Gingerich 1982; ⁶Hutchison 2013; ⁷Kondrashov and Lucas 2004; ⁸Meszoely et al. 1978; ⁹O'Leary and Rose 1995; ¹⁰Secord 2008; ¹¹Thewissen 1990