Grizzly Buttes (Eocene of the United States)

Where: Uinta County, Wyoming (41.2° N, 110.5° W: paleocoordinates 45.2° N, 96.8° W)

• coordinate based on political unit

• outcrop-level geographic resolution

When: lower Member (Bridger Formation), Bridgerian (50.3 - 46.2 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size classes: macrofossils, mesofossils

Collected by Marsh, Quigley, Harger, Matthew, Thomson, Gazin in 1870-1872, 1905; reposited in the AMNH, USNM, YPM

Primary reference: O. C. Marsh. 1872. Notice of some new Tertiary and post-Tertiary birds. The American Journal of Science and Arts, series 3 4(19-24):256-262 [M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 16228: authorized by John Alroy, entered by John Alroy on 26.03.1995, edited by Matthew Carrano, Philip Mannion, Roger Benson, Evangelos Vlachos, Patricia Holroyd and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

• mostly from Bridger B2 according to Matthew 1909 and Osborn 1929
type of "Triacodon nanus" is from Grizzly Buttes: Thorpe 1923a
Mammalia
 Soricomorpha - Geolabididae
"Nyctitherium dasypelix n. sp." = Marsholestes dasypelix4
"Nyctitherium dasypelix n. sp." = Marsholestes dasypelix4 Matthew 1909 placental
see Krishtalka 1976b; "found with that of Metacheiromys dasypus" (i.e., in Bridger B2): Simpson 1931
 Soricomorpha - Nyctitheriidae
Nyctitherium serotinum n. sp.4 Marsh 1872 placental
of Marsh see Matthew 1909 and Robinson 1968b
 Tribosphenida - Erinaceidae
Entomolestes grangeri n. gen. n. sp.4
Entomolestes grangeri n. gen. n. sp.4 Matthew 1909 hedgehog
from Bridger B2 at Grizzly Buttes West: see Matthew 1909 and Robinson 1968b
 Artiodactyla - Homacodontidae
Antiacodon pygmaeus28 Cope 1872 even-toed ungulate
Bridger B
 Artiodactyla - Helohyidae
Helohyus plicodon n. sp.4 Marsh 1872 even-toed ungulate
also at Bridger B2, see West 1984
 Perissodactyla - Equidae
Orohippus pumilus n. sp.4 Marsh 1871 horse
from "lower beds"; also "O. ballardi" type from Bridger B and referred specimen of "O. atavus": Granger 1908; see also Kitts 1957b
 Perissodactyla - Brontotheriidae
"Palaeosyops paludosus" = Palaeosyops paludosus23, "Palaeosyops major n. sp." = Palaeosyops paludosus16, "Limnohyops monoconus n. sp." = Palaeosyops paludosus4
"Palaeosyops paludosus" = Palaeosyops paludosus23 Leidy 1870 brontothere
from Bridger B2; one specimen is from Mountain View near Grizzly Buttes; also "P. major" type, from Grizzly Buttes (level unknown); type of "L. matthewi" from B2 at Grizzly Buttes West: see Osborn 1908, Osborn 1929
"Palaeosyops major n. sp." = Palaeosyops paludosus16 Leidy 1870 brontothere
"Limnohyops monoconus n. sp." = Palaeosyops paludosus4 Leidy 1870 brontothere
and "L. priscus" type Osborn 1908: both from B2 at Grizzly Buttes West also "cotype" of L. priscus from B2 at Grizzly Buttes East
 Perissodactyla - Hyrachyidae
Hyrachyus nanus16, "Hyrachyus modestus" = Hyrachyus modestus30, "Hyrachyus agrarius" = Hyrachyus modestus16, "Hyrachyus affinis" = Hyrachyus modestus30
Hyrachyus nanus16 Marsh 1871 odd-toed ungulate
"Hyrachyus modestus" = Hyrachyus modestus30 Leidy 1870 odd-toed ungulate
B2; also Grizzly Buttes East, B2
"Hyrachyus agrarius" = Hyrachyus modestus16 Leidy 1870 odd-toed ungulate
"Hyrachyus affinis" = Hyrachyus modestus30 Leidy 1870 odd-toed ungulate
B2
 Perissodactyla -
"Helaletes nanus n. sp." = Helaletes nanus4, "Helaletes boops n. gen. n. sp." = Helaletes nanus24
"Helaletes nanus n. sp." = Helaletes nanus4 Marsh 1871 odd-toed ungulate
see Radinsky 1963; see also Schoch 1984b
"Helaletes boops n. gen. n. sp." = Helaletes nanus24 Marsh 1871 odd-toed ungulate
 Ferae - Metacheiromyidae
Metacheiromys marshi n. sp.34 Wortman 1903 placental
referred material from B2, and type of "M. tatusia" from Bridger B2: see Simpson 1931, Schoch 1984a
Metacheiromys dasypus n. sp.4 Osborn 1904 placental
from Bridger B2: see Simpson 1931, Schoch 1984a
 Ferae - Viverravidae
Viverravus gracilis n. sp.4 Marsh 1872 placental
see Thorpe 1923a
Viverravus minutus32 Wortman 1901 placental
level not stated directly, but species is said to be restricted to the lower Bridger
 Ferae -
Miacis parvivorus19, "Miacis hargeri" = Vulpavus hargeri4, Vulpavus palustris n. sp.26, Vulpavus profectus n. sp.4, "Oodectes proximus n. sp." = Oodectes herpestoides4
Miacis parvivorus19 Cope 1872 placental
"Lower Bridger"
"Miacis hargeri" = Vulpavus hargeri4 Wortman 1901 placental
level not stated; see also Thorpe 1923a
Vulpavus palustris n. sp.26 Marsh 1871 placental
lower Bridger
Vulpavus profectus n. sp.4 Matthew 1909 placental
of Matthew 1909 from Bridger B2; see also Wang and Tedford 1994
"Oodectes proximus n. sp." = Oodectes herpestoides4 Wortman 1901 placental
of Matthew 1909 from "eastern part" of bluff
 Creodonta - Oxyaenidae
Patriofelis ulta26, "Patriofelis latidens n. sp." = Patriofelis ferox4
Patriofelis ulta26 Leidy 1870 creodont
i.d. based on size
"Patriofelis latidens n. sp." = Patriofelis ferox4 Marsh 1872 creodont
from the lower Bridger see Thorpe 1923a
 Creodonta -
"Limnocyon verus n. sp." = Limnocyon verus4, "Limnocyon riparius n. sp." = Limnocyon verus33, Thinocyon velox n. sp.4
"Limnocyon verus n. sp." = Limnocyon verus4 Marsh 1872 creodont
see Thorpe 1923a
"Limnocyon riparius n. sp." = Limnocyon verus33 Marsh 1872 creodont
Thinocyon velox n. sp.4 Marsh 1872 creodont
includes Marsh's referred specimen of "Triacodon fallax": Thorpe 1923a
 Creodonta - Hyaenodontidae
Sinopa rapax n. sp.4, Sinopa eximia n. sp.16, Tritemnodon agilis n. sp.4
Sinopa rapax n. sp.4 Leidy 1871 creodont
from Bridger B2 according to Matthew 1909; also "S. minor" type; also "Triacodon fallax" type of Thorpe 1923a
Sinopa eximia n. sp.16 Leidy 1873 creodont
Tritemnodon agilis n. sp.4 Marsh 1872 creodont
referred specimen discussed by Thorpe 1923a
 Acreodi - Mesonychidae
Mesonyx obtusidens26 Cope 1872 condylarth
 Primates - Microsyopidae
"Microsyops elegans n. sp." = Microsyops elegans34, "Microsyops gracilis" = Microsyops elegans16
"Microsyops elegans n. sp." = Microsyops elegans34 Marsh 1871 primate
see also Gunnell 1989
"Microsyops gracilis" = Microsyops elegans16 Marsh 1871 primate
 Primates - Notharctidae
"Notharctus tenebrosus" = Notharctus tenebrosus7, "Notharctus pugnax" = Notharctus tenebrosus8, Smilodectes gracilis n. sp.34
"Notharctus tenebrosus" = Notharctus tenebrosus7 Leidy 1870 primate
Bridger B; includes types of "Thinolestes anceps," "Limnotherium affine," "Notharctus formosus," and "N. osborni"; latter from B2 at Grizzly Buttes West, also referred "N. osborni" from B2 at Grizzly Buttes East and "N. tenebrosus" from B2 and B3, see Granger and Gregory 1917; also includes specimen of Hamrick and Alexander 1996 from "Black's Fork Member... Bridger B... 'Butch Hill' toward the northern end of Grizzly Buttes East, approximately 3 km southwest of the outlying butte Turtle Hill"
"Notharctus pugnax" = Notharctus tenebrosus8 Leidy 1870 primate
B2 at Grizzly Buttes East and B3 at Grizzly Buttes; confirmed by Gingerich 1979a
Smilodectes gracilis n. sp.34 Marsh 1871 primate
from Bridger B: see Gazin 1958a; also includes type of "Notharctus matthewi" from Bridger B2 at Grizzly Buttes East, see Granger and Gregory 1917, Gingerich 1979a; may include tentatively referred "Aphanolemur" of Granger and Gregory 1917 from lower Bridger at Grizzly Buttes West
 Primates - Omomyidae
Omomys carteri1 Leidy 1869 tarsier
Bridger B2, Grizzly Buttes East
Uintanius ameghini3 Wortman 1904 tarsier
"lower Bridger"; type of "U. turriculorum" of Matthew 1915c from "Horizon B2"; referred specimen from Bridger B, see Szalay 1976
 Rodentia - Reithroparamyidae
Reithroparamys delicatissimus n. sp.4 Leidy 1871 rodent
see Wood 1962
 Rodentia - Sciuravidae
? Tillomys parvidens n. sp.4 Marsh 1872 rodent
see Wilson 1938b
Sciuravus nitidus n. sp.4 Marsh 1871 rodent
see Wilson 1938a; includes type of "S. undans"
 Rodentia - Muridae
Mysops parvus n. sp.4, Mysops minimus n. sp.4
Mysops parvus n. sp.4 Marsh 1872 mouse
see Wilson 1938b
Mysops minimus n. sp.4 Leidy 1871 mouse
see Wilson 1938b
 Rodentia - Ischyromyidae
Leptotomus sp.12, Leptotomus parvus n. sp.4, "Thisbemys plicatus n. sp." = Thisbemys corrugatus4, Pseudotomus robustus n. sp.4, "Ischyrotomus horribilis n. sp." = Pseudotomus horribilis4, Paramys delicatus31
Leptotomus sp.12 Matthew 1910 rodent
size of L. bridgerensis; "L. grandis" Wood 1962 from "Bridger B of Grizzly Buttes West"
Leptotomus parvus n. sp.4 Wood 1959 rodent
from Bridger B2: see McGrew 1959, Wood 1962
"Thisbemys plicatus n. sp." = Thisbemys corrugatus4 Wood 1959 rodent
from Bridger B3: see Wood 1962
Pseudotomus robustus n. sp.4 Marsh 1872 rodent
see Wood 1962
"Ischyrotomus horribilis n. sp." = Pseudotomus horribilis4 Wood 1962 rodent
from Black's Fork Mbr.: see Wood 1962
Paramys delicatus31 Leidy 1871 rodent
Grizzly Buttes East, Bridger B3
 Condylarthra - Hyopsodontidae
Hyopsodus paulus19 Leidy 1870 condylarth
"lower beds" (Matthew 1909); "middle B" and "Grizzly Buttes West (middle B)" (West 1979); confirmed by Redline 1997
 Cimolesta - Esthonychidae
Trogosus castoridens n. sp.4 Leidy 1871 tillodont
includes "Palaeosyops minor" type, which also is from Grizzly Buttes, see Gazin 1953
 Cimolesta - Apatemyidae
Apatemys sp.27 Marsh 1872 eutherian
Reptilia
 Testudines - Pantrionychidae
"Plastomenus molopinus" = Anostira molopina10, "Aspideretes guttatus" = Trionyx guttatus10, "Platypeltis trionychoides" = Anostira trionychoides10, "Plastomenus thomasi" = Plastomenus thomasii10, Platypeltis trepida n. sp.9, "Platypeltis heteroglypta" = Trionyx heteroglyptus10
"Plastomenus molopinus" = Anostira molopina10 Cope 1872 turtle
AMNH 5941, 5945 from level B, collected 1903
"Aspideretes guttatus" = Trionyx guttatus10 Leidy 1869 turtle
AMNH 6005, 5917, 5993
"Platypeltis trionychoides" = Anostira trionychoides10 Cope 1872 turtle
AMNH 5938, 6017, 5942, found in 1903
"Plastomenus thomasi" = Plastomenus thomasii10 Cope 1872 turtle
AMNH 6018, nearly complete shell
Platypeltis trepida n. sp.9 Hay 1907 turtle
1905 AMNH collection from"Grizzly Buttes" B2
"Platypeltis heteroglypta" = Trionyx heteroglyptus10 Cope 1873 turtle
AMNH specimen, collected in 1903 from the eastern part of Grizzly Buttes, Wyoming
 Testudines - Carettochelyidae
Anosteira ornata10 Leidy 1871 turtle
some of the four, now lost, types (possibly including USNM 4062), others were from Church Buttes [PaleoDB collection 16201]
 Testudines - Geoemydidae
"Echmatemys pusilla n. sp." = Bridgeremys pusilla11, "Clemmys morrisiae n. sp." = Bridgeremys pusilla10, Echmatemys arethusa n. sp.10, "Echmatemys aegle n. sp." = Echmatemys stevensoniana10, "Echmatemys cyane n. sp." = Echmatemys stevensoniana10
"Echmatemys pusilla n. sp." = Bridgeremys pusilla11 Hay 1908 turtle
AMNH 5992 - type (partial shell)
"Clemmys morrisiae n. sp." = Bridgeremys pusilla10 Hay 1908 turtle
AMNH 6029 (holotype), a partial shell (Hay 1908, figs. 359, 360, pl. 45.1–3)
Echmatemys arethusa n. sp.10 Hay 1908 turtle
AMNH 5920 (holotype), a partial shell (Hay 1908, figs. 421, 422, pl. 49)
"Echmatemys aegle n. sp." = Echmatemys stevensoniana10 Leidy 1870 turtle
AMNH 5909 (holotype), a complete plastron and some carapacial bones (Hay 1908, figs. 436–438)
"Echmatemys cyane n. sp." = Echmatemys stevensoniana10 Leidy 1870 turtle
AMNH 5924 (holotype), a partial shell (Hay 1908, figs. 423–427, pl. 49)
 Testudines - Pantestudinidae
"Testudo corsoni n. sp." = Hadrianus corsoni15
"Testudo corsoni n. sp." = Hadrianus corsoni15 Leidy 1871 turtle
 Testudines - Trionychidae
"Aspideretes ellipticus" = Trionyx ellipticus10, "Amyda mira n. sp." = Trionyx mira10, "Platypeltis extensa n. sp." = Trionyx mira10, "Platypeltis postera" = Trionyx mira10, "Amyda franciscae n. sp." = Trionyx uintaensis10, Amyda exquisita n. sp.10
"Aspideretes ellipticus" = Trionyx ellipticus10 Hay 1908 softshell turtle
AMNH 5930
"Amyda mira n. sp." = Trionyx mira10 Hay 1908 softshell turtle
AMNH 6130, holotype partial skeleton discovered by the American Museum party of 1905, at Grizzly Buttes, Wyoming, in level B, of the Bridger Eocene
"Platypeltis extensa n. sp." = Trionyx mira10 Hay 1908 softshell turtle
AMNH 5951, holotype partial shell collected by O. P. Hay in 1903, western part of Grizzly Buttes
"Platypeltis postera" = Trionyx mira10 Hay 1908 softshell turtle
AMNH 5955, shell collected 1906
"Amyda franciscae n. sp." = Trionyx uintaensis10 Cope 1873 softshell turtle
AMNH 5936 (holotype) partial skeleton, level B
Amyda exquisita n. sp.10 Hay 1908 softshell turtle
portions of 2 individuals, which were collected in the Bridger beds of the Grizzly Buttes, Wyoming, by the museum's expedition of 1903. Of these specimens, number 5923 is taken as the type. It consists of the carapace (fragmentary but representing well the structure), the hyoplastra and hypoplastra of both sides somewhat damaged, one cervical vertebra, portions of both humeri, the left femur, portions of the pelvis, and various other leg and foot bones. Of the other individual, No. 5943, there are present only the hyoplastron and the hypoplastron of the right side
 Testudinata - Baenidae
"Baena riparia n. sp." = Baena affinis10, "Baena arenosa" = Baena arenosa10, "Baena sima" = Baena arenosa10, Chisternon undatum10, Chisternon undatum2
"Baena riparia n. sp." = Baena affinis10 Leidy 1871 turtle
AMNH 5977 (type of Baena riparia Hay, 1908, figs. 59-66, pls. 17, 18, 19 figs. 1-3), an almost complete skeleton
"Baena arenosa" = Baena arenosa10 Leidy 1870 turtle
AMNH 5900, AMNH 5906, AMNH 5921, AMNH 5922, AMNH 5935, AMNH 5957, AMNH 5965, AMNH 5985, AMNH 5986, AMNH 5990, all shells; AMNH 5952, shell, skull, limbs; AMNH 5973 (Hay, 1908, figs. 44-48, pl. 12), skull, shell, postcranium; AMNH 5984, shell, lower jaws, limb; USNM 18102, skull
"Baena sima" = Baena arenosa10 Leidy 1870 turtle
AMNH 5965, partial shell; 5907, partial skeleton with shell (informal paratypes)
Chisternon undatum10 Leidy 1871 turtle
AMNH 5959 (Hay, 1908, p. 86, fig. 72), a partial shell; AMNH 5962 (Hay, 1908, pp. 86, 87, figs. 73- 75), shell, skull, vertebrae, limb elements; AMNH 5964, shell. Locality: Grizzly Buttes West, Bridger Basin, Wyoming (label) (see Wheeler, 1961, pl. 2)
Chisternon undatum2 Leidy 1871 turtle
AMNH 5904, a shell, skull, vertebrae, and limb elements (referred to Chisternon hebraicum by Hay, 1908, pp. 89-91, figs. 82-87); AMNH 5961 (referred to Chisternon hebraicum by Hay, 1908, pp. 87-89, figs. 76-81, pl. 21 figs. 3, 4, pl. 23 fig. 1)
Aves
 Strigiformes - Protostrigidae
Protostrix leptosteus29 Marsh 1871 owl
YPM type specimen of Marsh collected in 1870 from "Grizzly Buttes"; Rich 1982 describes an additional AMNH specimen from the lower Bridger at "Grizzly Buttes" collected in 1903 and called a "falconine" by Shufeldt 1913
 Gruiformes - Gruidae
Aletornis nobilis n. gen. n. sp., Aletornis bellus n. sp.
Aletornis nobilis n. gen. n. sp. Marsh 1872 crane
Aletornis bellus n. sp. Marsh 1872 crane
Reptilia
 Crocodylia - Planocraniidae
"Pristichampsus vorax" = Boverisuchus vorax14
"Pristichampsus vorax" = Boverisuchus vorax14 Troxell 1925 crocodilian
YPM 5890 (incomplete right quadrate)
 Crocodylia - Crocodylidae
Crocodilus ziphodon n. sp.17, "Crocodylus affinis n. sp." = Crocodylus affinis22, "Brachyuranochampsa zangerli n. sp." = Crocodylus affinis21, "Crocodilus liodon n. sp." = Crocodylus affinis17, "Crocodilus brevicollis n. sp." = Crocodylus affinis22
Crocodilus ziphodon n. sp.17 Marsh 1871 crocodile
YPM 1347
"Crocodylus affinis n. sp." = Crocodylus affinis22 Marsh 1871 crocodile
YPM 1345 - type (skull and mandible)
"Brachyuranochampsa zangerli n. sp." = Crocodylus affinis21 Marsh 1871 crocodile
1903 AMNH Thomson collection from "Grizzly Buttes"
"Crocodilus liodon n. sp." = Crocodylus affinis17 Marsh 1871 crocodile
YPM 1350b - type (very fragmentary skeleton including several vertebrae, the left quadrate, rostral fragments, and articulars and surangulars)
"Crocodilus brevicollis n. sp." = Crocodylus affinis22 Marsh 1871 crocodile
YPM 1349 - type (three cervical vertebrae)
 Crocodylia - Alligatoridae
Allognathosuchus polyodon20 Cope 1873 crocodilian
L. S. Quackenbush AMNH collection from "Grizzly Buttes"
 Squamata - Rhineuridae
"Glyptosaurus anceps n. sp." = Lestophis anceps5
"Glyptosaurus anceps n. sp." = Lestophis anceps5 Marsh 1871 worm lizard
YPM 520 (holotype), single dorsal vertebra
 Squamata -
"Boavus occidentalis n. gen. n. sp." = Boavus occidentalis6, "Boavus agilis n. sp." = Boavus occidentalis6, Boavus brevis n. sp.6
"Boavus occidentalis n. gen. n. sp." = Boavus occidentalis6 Marsh 1871 snake
PMNH 511 (holotype), single dorsal vertebra; PMNH 2717, single dorsal vertebra
"Boavus agilis n. sp." = Boavus occidentalis6 Marsh 1871 snake
PMNH 2765 and 2766 (cotypes), thoracic vertebrae
Boavus brevis n. sp.6 Marsh 1871 snake
PMNH 468 (holotype), single thoracic vertebra
 Squamata - Boidae
Lithophis sargenti n. gen. n. sp.6
Lithophis sargenti n. gen. n. sp.6 Marsh 1871 boa
YPM 2719 (holotype), single thoracic vertebra
 Squamata -
"Thinosaurus leptodus n. sp." = Saniwa ensidens18, "Thinosaurus grandis n. sp." = Saniwa ensidens18, "Thinosaurus paucidens n. gen. n. sp." = Saniwa ensidens18
"Thinosaurus leptodus n. sp." = Saniwa ensidens18 Leidy 1870 squamates
Originally Thinosaurus specimens: YPM 612 (holotype), anterior portion of right dentary, plus posterior skeleton; YPM 613, partial postcranial skeleton. Additional Saniwa specimens: YPM 1064, 10 dorsal vertebrae
"Thinosaurus grandis n. sp." = Saniwa ensidens18 Leidy 1870 squamates
YPM 611 (part) (holotype), trunk vertebra lacking the condylar end
"Thinosaurus paucidens n. gen. n. sp." = Saniwa ensidens18 Leidy 1870 squamates
YPM (unknown, lost holotype), greater part of a skeleton
 Squamata - Anguidae
Oreosaurus vagans n. gen. n. sp.18, "Glyptosaurus sylvestris n. gen. n. sp." = Glyptosaurus sylvestris5, "cf. Glyptosaurus sylvestris" = Glyptosaurus sylvestris25, "Glyptosaurus princeps n. sp." = Glyptosaurus sylvestris18, "Paraglyptosaurus princeps" = Glyptosaurus sylvestris25, "cf. Paraglyptosaurus princeps" = Glyptosaurus sylvestris25, "Glyptosaurus nodosus n. sp." = Glyptosaurus sylvestris5, "Glyptosaurus brevidens n. sp." = Glyptosaurus sylvestris18, "Glyptosaurus rugosus n. sp." = Glyptosaurus sylvestris18
Oreosaurus vagans n. gen. n. sp.18 Marsh 1872 squamates
YPM 541 (holotype), frontals, portions of dentaries, partial coronoid, fragment of pterygoid with teeth. numerous dermal scutes, other fragments of skull and jaws
"Glyptosaurus sylvestris n. gen. n. sp." = Glyptosaurus sylvestris5 Marsh 1871 squamates
YPM 526 (=USNM 6523) (holotype), left frontal; YPM 523 (prev. holotype of G. ocellatus), small patch of cranial scutes; YPM 521 (prev. holotype of G. brevidens), many parts of broken skull and jaws, several short articulated series of cervical and dorsal vertebrae and numerous skull and body osteoderm patches; YPM 1073, partial skull and postcranials; AMNH 6054 now referred to P. princeps (ref 16228); AMNH 5113, partial skull referred to G. princeps by Gilmore 1928, to G. sylv. by Sullivan 1979; AMNH 5902, partial skull
"cf. Glyptosaurus sylvestris" = Glyptosaurus sylvestris25 Marsh 1871 squamates
YPM 1073, fragment of left frontal
"Glyptosaurus princeps n. sp." = Glyptosaurus sylvestris18 Marsh 1871 squamates
YPM 524 (=USNM 16539) (holotype), considerable portion of coalesced frontals, median part of right pterygoid, fragm. portions of both rami including symph. end of left dentary, dermal scutes, phalangeal bone, other fragments; (AMNH 5113 now G. sylvestris); AMNH 6054, nearly complete parietal, portion of left jugal, and alveolar border of one maxilla with six teeth, plus osteoderms (prev. G. sylvestris)
"Paraglyptosaurus princeps" = Glyptosaurus sylvestris25 Marsh 1871 squamates
YPM 1072, partial skull and postcranials (previously ID'd as G. sylvestris)
"cf. Paraglyptosaurus princeps" = Glyptosaurus sylvestris25 Marsh 1871 squamates
YPM 7581, frontal fragment
"Glyptosaurus nodosus n. sp." = Glyptosaurus sylvestris5 Marsh 1871 squamates
YPM 522 (=USNM 16520) (holotype), frontal, two body scutes, unidentified skull fragment
"Glyptosaurus brevidens n. sp." = Glyptosaurus sylvestris18 Marsh 1871 squamates
"Glyptosaurus rugosus n. sp." = Glyptosaurus sylvestris18 Marsh 1871 squamates
YPM 525 (= USNM 16526) (holotype), posterior portions of coosified frontal bones
Baptemys wyomingensis
  -
Baptemys wyomingensis10 Leidy 1870
AMNH 5934 (shell), 5967 (shell and large portion of the skull)
Scenopagus edenensis
  -
Scenopagus edenensis13 McGrew et al. 1959
lower Bridger