Amöneburg (Miocene of Germany)

Also known as Amoneburg

Where: Rhineland-Palatinate, Germany (50.0° N, 8.3° E: paleocoordinates 50.1° N, 6.2° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wiesbaden Formation, MN 2 (22.4 - 20.0 Ma)

• "The Formation [i.e. the Wiesbaden Formation, formerly referred to as the Lower Hydrobia beds; TL] can be divided into three parts. The lower (brackish) and middle (lacustrine) parts are middle Aquitanian, containing the MN 2 mammal zone (Tobien 1988; Gers & Medden 1991; Reichenbacher 2000; Reichenbacher & Keller 2002). Reichenbacher (2000) assumed that the MN 2b and MN 3a mammal sub-zones of the Late Aquitanian–Early Burdigalian interval corresponded to the upper part of the Wiesbaden Formation, named the ‘emerging facies’. The squamate microvertebrate material described here was collected by Thomas Keller and colleagues from the lower brackish part of the Wiesbaden Formation where numerous freshwater and terrestrial gastropods, including Gyraulus, Stagnicola and Planorbarius, also occur." Age of the lower part of the Wiesbaden Fm is given as MN2a.

Environment/lithology: lagoonal or restricted shallow subtidal; limestone

• No lithology description is given in the primary reference. The Wiesbaden Formation, however, belongs to the so-called "Kalktertiär" ("limestone Tertiary") of the Mainz Basin. Hence, the host rocks may well have been calcareous rocks.

Size class: mesofossils

Collected by Thomas Keller et al.; reposited in the SMF

Collection methods: sieve,

• "The fossils were recovered by screen washing and are housed in the SMF, prefixed by collection numbers of the PAL EV."

Primary reference: A. Cernansky, J. C. Rage, and J. Klembara. 2015. The Early Miocene squamates of Amöneburg (Germany): the first stages of modern squamates in Europe. Journal of Systematic Palaeontology 13(2):97-128 [J. Mueller/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 167464: authorized by Johannes Mueller, entered by Torsten Liebrecht on 18.03.2015

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Squamata -
Squamata indet. squamates
PAL EV 2013/272 (atlas fragemnt); PAL EV 2013/273 and 274 (fragments of humeri: two proximal and one distal part); PAL EV 2013/275 (1 left pelvis)
Merkurosaurus ornatus Klembara 2008 squamates
PAL EV 2013/268 (a jugal)
cf. Merkurosaurus ornatus Klembara 2008 squamates
PAL EV 2013/277 (a single osteoderm)
Platynota indet. Baur 1890 squamates
PAL EV 2013/270 (a single osteoderm)
Shinisauria indet. Conrad 2008 squamates
PAL EV 2013/269 (fragment of a parietal)
 Squamata - Anguidae
Anguinae indet. Gray 1825 squamates
PAL EV 2013/176 (a right quadrate); PAL EV 2013/177 to 179 (3 dorsal vertebrae); PAL EV 2013/180 and 181 (2 caudal vertebrae); PAL EV 2013/182 to 267 (86 osteoderms)
Pseudopus sp. Merrem 1820 legless lizard
PAL EV 2013/166 to 170 (5 osteoderms)
Pseudopus cf. ahnikoviensis Klembara 2012 legless lizard
PAL EV 2013/165 (a right dentary)
Ophisaurus sp. Daudin 1803 glass lizard
PAL EV 2013/172 to 174 (3 frontals: 1 left, 2 right); PAL EV 2013/175 (a right pterygoid); PAL EV 2013/276 (a right dentary)
Ophisaurus spinari Klembara 1979 glass lizard
PAL EV 2013/171 (a parietal)
 Squamata -
"Scincomorpha" indet. Camp 1923 squamates
PAL EV 2013/164 (a right dentary)
Gekkota indet. Cuvier 1817 squamates
PAL EV 2013/147 (left maxilla)
 Squamata - Lacertidae
Lacertidae indet. Bonaparte 1831 squamates
PAL EV 2013/155 (fragment of a right frontal); PAL EV 2013/156 to 159 (4 jugals: 2 left, 2 right); PAL EV 2013/160 (a right pterygoid); PAL EV 2013/161 (a basisphenoid); PAL EV 2013/162 to 163 (2 dorsal vertebrae)
Lacerta poncenatensis Müller 1996 squamates
PAL EV 2013/148 to 151 (4 dentary fragments: 2 of a left, 2 of a right dentary); PAL EV 2013/152 and 153 (2 maxilla fragments: 1 of a left, 1 of a right maxilla); PAL EV 2013/154 (a premaxilla)
 Squamata -
Serpentes indet. Linnaeus 1758 snake
PAL EV 2013/111 (an incomplete compound bone); PAL EV 2013/112 to 146 (35 trunk and caudal vertebrae)
 Squamata - Colubridae
cf. Neonatrix sp. Holman 1973 colubrid snake
PAL EV 2013/11 and 12 (2 trunk vertebrae: 1 anterior, 1 posterior)
? Neonatrix sp. Holman 1973 colubrid snake
PAL EV 2013/110 (a single cloacal vertebra)
Coluber "sp. 1" Linnaeus 1758 racer
"Coluber sensu lato"; PAL EV 2013/13 to 84 (72 trunk vertebrae)
Coluber "sp. 2" Linnaeus 1758 racer
"Coluber sensu lato"; PAL EV 2013/85 to 104 (20 trunk vertebrae)
Coluber indet. Linnaeus 1758 racer
"Coluber sensu lato"; PAL EV 2013/105 (incomplete parabasisphenoid)
cf. Natrix sp. Laurenti 1768 water snake
PAL EV 2013/8–10 (3 trunk vertebrae)
 Squamata -
Booidea indet. Gray 1825 snake
PAL EV 2013/6 and 7 (2 trunk vertebrae)
 Squamata - Boidae
? Falseryx sp. Szyndlar and Rage 2003 boa
PAL EV 2013/2 to 5 (4 trunk vertebrae)
 Squamata -
Colubroidea indet. Oppel 1811 snake
PAL EV 2013/106 (a single trunk vertebra)
 Squamata - Crotalidae
Vipera sp. Laurenti 1768 pit viper
"aspis complex"; PAL EV 2013/107 to 109 (3 trunk vertebrae)
 Squamata - Amphisbaenidae
Blanus thomaskelleri n. sp. Cernansky et al. 2015 worm lizard
PAL EV 2013/271 (holotype: incomplete right dentary)