Mustersan of Patagonia (Eocene of Argentina)

Where: Chubut, Argentina (45.7° S, 68.7° W: paleocoordinates 48.1° S, 58.8° W)

• coordinate based on political unit

• basin-level geographic resolution

When: Mustersan mammal zone, Mustersan (48.0 - 42.0 Ma)

• Mustersan SALMA, Late Eocene (Woodburne et al. 2014).

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size classes: macrofossils, mesofossils

Collected by Carlos Ameghino in c. 1900

Primary reference: G. G. Simpson. 1948. The beginning of the age of mammals in South America. Part I. Bulletin of the American Museum of Natural History 91:1-232 [D. Croft/D. Croft/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 176286: authorized by Jelle Zijlstra, entered by Jelle Zijlstra on 06.02.2016

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Theriamorpha - Pyrotheriidae
Propyrotherium saxeum n. gen. n. sp.
Propyrotherium saxeum n. gen. n. sp. Ameghino 1901 placental
MACN 10929, ?P4 (lectotype)
 Notoungulata -
Ultrapithecus robustus n. sp. Roth 1902 notoungulate
Type not relocated by Simpson (1967), said to be "[p]resumably Mustersan"
Procolpodon foratus n. gen. n. sp. Ameghino 1904 notoungulate
Type not relocated by Simpson (1967), left astragalus
 Notoungulata - Archaeohyracidae
Eohyrax brachyodus n. sp., Eohyrax platyodus n. sp., "Archaeohyrax sulcidens n. sp." = Bryanpattersonia sulcidens, "Archaeohyrax nesodontoides n. sp." = Bryanpattersonia nesodontoides, "Eomorphippus rutilatus n. sp." = Bryanpattersonia nesodontoides, Pseudhyrax eutrachytheroides n. gen. n. sp., "Eohyrax strangulatus n. sp." = Pseudhyrax strangulatus, "Trimerostephanos cuneolus n. sp." = Pseudhyrax strangulatus
Eohyrax brachyodus n. sp. Ameghino 1902 notoungulate
Type not relocated, probably an isolated m1
Eohyrax platyodus n. sp. Ameghino 1904 notoungulate
MACN 10980, six teeth of which two are referable to this species: left m1 (lectotype) and left m3
"Archaeohyrax sulcidens n. sp." = Bryanpattersonia sulcidens Ameghino 1902 notoungulate
MACN 1096, many isolated teeth, including left m1, lectotype
"Archaeohyrax nesodontoides n. sp." = Bryanpattersonia nesodontoides Ameghino 1901 notoungulate
MACN 10905, 10 lower and 7 upper teeth, including right m1, lectotype
"Eomorphippus rutilatus n. sp." = Bryanpattersonia nesodontoides Ameghino 1901 notoungulate
MACN 10915, a right upper premolar (doubtfully referable), two left, and one right (lectotype) upper molars
Pseudhyrax eutrachytheroides n. gen. n. sp. Ameghino 1901 notoungulate
MACN, uncatalogued, maxillary fragment with dP2-4, M1
"Eohyrax strangulatus n. sp." = Pseudhyrax strangulatus Ameghino 1901 notoungulate
MACN 10774, dentary p4-m2, type (but not other specimens cataloged under this number); MACN 10913, dentary with partial p4 and m1-2
"Trimerostephanos cuneolus n. sp." = Pseudhyrax strangulatus Ameghino 1901 notoungulate
MACN 10968, six associated lower cheek teeth and jaw fragments
 Notoungulata - Interatheriidae
Eopachyrucos pliciformis n. gen. n. sp.
Eopachyrucos pliciformis n. gen. n. sp. Simpson 1967 notoungulate
MACN A55-12, isolated upper molar
 Notoungulata -
"Eomorphippus obscurus n. gen. n. sp." = Eomorphippus obscurus, "Pseudostylops subquadratus n. gen. n. sp." = Eomorphippus obscurus, "Proasmodeus exauctus n. sp." = Periphragnis exauctus, "Asmodeus armatus n. sp." = Periphragnis harmeri
"Eomorphippus obscurus n. gen. n. sp." = Eomorphippus obscurus Ameghino 1901 notoungulate
MACN 10917, three upper molars (including left M3, lectotype); MACN 10914, isolated teeth including 7 referable to this species. Perhaps from the "Astraponotéen plus supérieure" (=Tinguirirican) level but no concrete evidence of that for these specimens.
"Pseudostylops subquadratus n. gen. n. sp." = Eomorphippus obscurus Ameghino 1901 notoungulate
MACN 10904, P?3
"Proasmodeus exauctus n. sp." = Periphragnis exauctus Ameghino 1902 notoungulate
MACN 10926, mandible p2-m3 (type); doubtfully referred MACN 10928, mandible right p2-m3, left m2-3
"Asmodeus armatus n. sp." = Periphragnis harmeri Roth 1899 notoungulate
MACN 10925, P2-M3 on both sides, lectotype. See discussion of original hypodigm by Simpson (1967).
 Notoungulata - Isotemnidae
Trimerostephanos coalitus n. sp., Trimerostephanos coarctatus n. sp., "Trimerostephanos sigma n. sp." = Acoelohyrax sigma
Trimerostephanos coalitus n. sp. Ameghino 1901 notoungulate
MACN 10964, right dentary with ?p4-m1
Trimerostephanos coarctatus n. sp. Ameghino 1901 notoungulate
MACN 10965A, dentary p4-m3, lectotype; MACN 10965B, right P2-4, M2-3, left P2-4, M2-3; MACN 10966B, M?2; other specimens under MACN 10965-10966 were referred by Ameghino but may not belong here
"Trimerostephanos sigma n. sp." = Acoelohyrax sigma Ameghino 1901 notoungulate
MACN 10967, mandible m1-2, type; MACN 10969, dentary m1-3
 Notoungulata - Notopithecidae
"Notopithecinae indet." = Notopithecidae, "Guilielmoscottia plicifera n. gen. n. sp." = Guilielmoscottia plicifera, "Adpithecus amplidens n. sp." = Guilielmoscottia plicifera
"Notopithecinae indet." = Notopithecidae Ameghino 1897 notoungulate
? Notopithecus amplidens or Guilielmoscottia plicifera. MACN 10786, dentary p1-m3
"Guilielmoscottia plicifera n. gen. n. sp." = Guilielmoscottia plicifera Ameghino 1901 notoungulate
MACN 10898, maxilla P1-M3
"Adpithecus amplidens n. sp." = Guilielmoscottia plicifera Ameghino 1901 notoungulate
MACN 10895, maxilla P3-M3 (lectotype) and maxilla dP2-4, M1-2; MACN 10901, C-P2, P4-M2, m3, and upper premolar
 Notoungulata -
Edvardocopeia sinuosa n. gen. n. sp. Ameghino 1901 notoungulate
Type not relocated by Simpson (1967), but original description based on an upper premolar and a symphysis
Pseudopachyrucos foliiformis n. gen. n. sp. Ameghino 1901 notoungulate
MACN A55-11, three isolated, broken teeth, including the lectotype
Eohegetotherium priscum n. gen. n. sp. Ameghino 1901 notoungulate
MACN, uncatalogued lot, isolated upper molar (lectotype)
Trimerostephanos ultimus n. sp. Ameghino 1901 notoungulate
Type not relocated by Simpson (1967), but based on m1 and M1-3
 Panameriungulata -
Paulogervaisia celata n. sp. Ameghino 1901 placental
MACN 10947, tooth
 Panameriungulata - Macraucheniidae
"Lambdaconus alius n. sp." = Polymorphis alius
"Lambdaconus alius n. sp." = Polymorphis alius Ameghino 1901 placental
MACN 10893, mandible with left p2-m1, right p2-m3
 Panameriungulata - Proterotheriidae
"Decaconus intricatus n. gen. n. sp." = Polyacrodon lanciformis
"Decaconus intricatus n. gen. n. sp." = Polyacrodon lanciformis Roth 1899 placental
MACN 10897, right upper molar
 Cingulata - Glyptatelidae
Lomaphorelus depstus n. gen. n. sp. Ameghino 1902 edentate
MACN 10950, broken scute
Glyptatelus fractus n. sp. Ameghino 1902 edentate
MACN 10949, scute (lectotype) and isolated tooth
 Cingulata - Palaeopeltidae
Palaeopeltis tesseratus n. sp. Ameghino 1902 edentate
MACN 10948, broken fragment of shield with most of one scute and small parts of three others
 Cingulata - Dasypodidae
"Anteutatus laevus n. sp." = Utaetus laevus, Machlydotherium ater n. sp., Machlydotherium intortum n. sp., Machlydotherium asperum n. gen. n. sp., "Meteutatus rigidus n. sp." = Barrancatatus rigidus, Sadypus ascendens n. sp., Sadypus confluens n. sp., "Anutaetus circundatus n. gen. n. sp." = Pseudeutatus circundatus, "Isutaetus depictus n. gen. n. sp." = Pseudeutatus depictus, "Pachyzaedyus cuneiformis n. gen. n. sp." = Pseudeutatus cuneiformis, Pseudeutatus clypeus n. gen. n. sp.
"Anteutatus laevus n. sp." = Utaetus laevus Ameghino 1902 armadillo
MACN 10690, six scutes
Machlydotherium ater n. sp. Ameghino 1902 armadillo
MACN 10952, ten scutes
Machlydotherium intortum n. sp. Ameghino 1902 armadillo
MACN 10953, three scutes
Machlydotherium asperum n. gen. n. sp. Ameghino 1902 armadillo
MACN 10951, 28 scutes, many broken, and one tooth
"Meteutatus rigidus n. sp." = Barrancatatus rigidus Ameghino 1902 armadillo
MACN 10958, twelve scutes
Sadypus ascendens n. sp. Ameghino 1902 armadillo
MACN 10955, one scute
Sadypus confluens n. sp. Ameghino 1902 armadillo
MACN 10954, eight scutes
"Anutaetus circundatus n. gen. n. sp." = Pseudeutatus circundatus Ameghino 1902 armadillo
MACN 10962, one scute
"Isutaetus depictus n. gen. n. sp." = Pseudeutatus depictus Ameghino 1902 armadillo
MACN 10961, one band and two buckler scutes
"Pachyzaedyus cuneiformis n. gen. n. sp." = Pseudeutatus cuneiformis Ameghino 1902 armadillo
MACN 10956, one scute
Pseudeutatus clypeus n. gen. n. sp. Ameghino 1902 armadillo
MACN 10959, one scute
 Sparassodonta -
Procladosictis anomala n. gen. n. sp.
Procladosictis anomala n. gen. n. sp. Ameghino 1902 metatherian
MACN 10327, maxilla P3-M3, holotype
 Theriamorpha -
? Promysops primarius n. sp. Ameghino 1902 marsupial
MACN A55-10, isolated incisor, thought to be lost by Simpson (1948) but relocated by Simpson (1967).
  -
Picunia nitida n. gen. n. sp., Diplodon ampliatus n. gen. n. sp., Stenogenium aenigmaticum n. sp.
Picunia nitida n. gen. n. sp. Roth 1902 mammal
"Probably an isolated upper molar; not found in collection"
Diplodon ampliatus n. gen. n. sp. Roth 1902 mammal
Isolated tooth, not relocated by Simpson (1967)
Stenogenium aenigmaticum n. sp. Ameghino 1901 mammal
MACN 10899, edentulous symphysis