Puesto Estancia La Costa Fossiliferous Level 5.3 (Miocene of Argentina)

Where: Santa Cruz, Argentina (51.2° S, 69.1° W: paleocoordinates 51.8° S, 63.4° W)

• coordinate stated in text

• local area-level geographic resolution

When: Estancia La Costa Member (Santa Cruz Formation), Santacrucian (17.5 - 16.3 Ma)

• The age of this level is between 17.3 and 17.6 Ma, based on the age of two radiometrically dated tuffs that bracket it: the CO tuff of Perkins et al. (2012) and the CV-13 tuff of Trayler et al. (2020). The former as refined by high-precision U–Pb age modeling is 17.31 Ma and the latter is 17.62 Ma (Trayler et al., 2020).

• formation-level stratigraphic resolution

Environment/lithology: terrestrial; green, sandy claystone and tuffaceous, silty sandstone

Size class: macrofossils

Primary reference: F. J. Degrange, J. I. Noriega, and J. I. Areta. 2012. Diversity and paleobiology of the Santacrucian birds. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 138-155 [J. Zijlstra/J. Zijlstra]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 176602: authorized by Jelle Zijlstra, entered by Jelle Zijlstra on 17.02.2016

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Megatherioidea - Megalonychidae
cf. Hyperleptus sp.2 Ameghino 1891 edentate
MPM-PV 3410, skull and postcrania
cf. Hapalops sp.2 Ameghino 1887 edentate
MPM-PV 3462, incomplete skull and postcrania
Hapalops sp.2 Ameghino 1887 edentate
MPM-PV 3412, skull and postcrania; MPM-PV 3467, skull and postcrania
Eucholoeops ingens2 Ameghino 1887 edentate
MPM-PV 3451, skull and postcrania. See also ref 71644
  - Mylodontidae
Nematherium sp.2 Ameghino 1887 edentate
MPM-PV 3703, mandible
 Cingulata - Chlamyphoridae
Prozaedyus proximus7 Ameghino 1887 edentate
MPM-PV 3418, skull fragment; MPM-PV 3423, skull; MPM-PV 3506, complete skeleton; MPM-PV 3617, skeleton and carapace; MPM-PV 4182, skull, mandible, osteoderms
"Proeutatus oenophorus" = Proeutatus oenophorum7 Ameghino 1887 edentate
MPM-PV 3415, skull and postcrania; MPM-PV 3425, skull fragment; MPM-PV 3430, skull and postcrania; MPM-PV 4256, skull
Stenotatus cf. patagonicus7 Ameghino 1887 edentate
MPM-PV 3429, mandible and osteoderms
 Cingulata - Peltephilidae
Peltephilus pumilus7 Ameghino 1887 edentate
MPM-PV 3414, skull and pelvis fragment; MPM-PV 3431, osteoderms
 Cingulata - Propalaehoplophoridae
Cochlops muricatus7 Ameghino 1889 edentate
MPM-PV 3420, skull, partial skeleton, carapace
Propalaehoplophorus australis7 Moreno 1882 edentate
MPM-PV 3422, skull and postcrania
 Notoungulata - Interatheriidae
"Protypotherium praerutilum" = Protypotherium praerutilum4, "Protypotherium attenuatum" = Protypotherium praerutilum4
"Protypotherium praerutilum" = Protypotherium praerutilum4 Ameghino 1887 notoungulate
MPM-PV 3530, mandible; 3659, cranium and mandible;
"Protypotherium attenuatum" = Protypotherium praerutilum4 Ameghino 1887 notoungulate
MPM-PV 3470, cranium and mandible
 Notoungulata - Hegetotheriidae
Hegetotherium mirabile4 Ameghino 1887 notoungulate
MPM-PV 3526, cranium; 4316, mandible
 Notoungulata - Toxodontidae
Nesodon imbricatus4 Owen 1846 notoungulate
MPM-PV 4377, cranium
 Rodentia -
Neoreomys australis3 Ameghino 1887 caviomorph
MPM-PV 3514, partial skull, left femur; 3674, skull, postcrania; 3676, partial skull; 4340, skull and postcrania; 4341, skull and postcrania; 4342, skull and postcrania; 4343, skull and vertebrae; 4344, dentary
Steiromys detentus3 Ameghino 1887 caviomorph
MPM-PV 4358, dentary dp4–m2; 4271, partial skull
Steiromys duplicatus3 Ameghino 1887 caviomorph
MPM-PV 4273, skull and right mandible
 Primates - Homunculidae
Homunculus patagonicus5 Ameghino 1891 monkey
MPM-PV 3500, humerus (see also Kay et al. 2021); 3501, cranium; 3503, cranium; 3504, mandible; 3505, juvenile cranium (see ref 58229)
 Panameriungulata - Proterotheriidae
Thoatherium minusculum4 Ameghino 1887 placental
MPM-PV 3529, cranium, femur, ulna, tibia
Tetramerorhinus cingulatum4 Ameghino 1891 placental
MPM-PV 3493, cranium
 Paucituberculata - Palaeothentidae
Palaeothentes lemoinei1 Ameghino 1887 marsupial
MPM-PV 3494, dentary m2-4, humerus, ulna, pelvis, tibia, astragalus
 Sparassodonta - Borhyaenidae
Prothylacynus patagonicus6 Ameghino 1891 metatherian
MPM-PV 4318, 4331
Borhyaena tuberata6 Ameghino 1887 metatherian
MPM-PV 3625, 4254, 4321, 4320
 Sparassodonta -
Sipalocyon sp.6 Ameghino 1887 metatherian
MPM-PV 4319
Sipalocyon gracilis6 Ameghino 1887 metatherian
MPM-PV 4317
? Perathereutes pungens6 Ameghino 1891 metatherian
MPM-PV 4322
Cladosictis patagonica6 Ameghino 1887 metatherian
MPM-PV 4253
Aves
 Gruiformes - Phorusrhacidae
Psilopterus bachmanni Moreno and Mercerat 1891 terror bird
MPM-PV 4243, furcula and left fore- and hindlimb fragments
 Cariamiformes - Cariamidae
Cariama santacrucensis n. sp. Noriega et al. 2009 seriema
MPM-PV 3511, incomplete cranium
Cariaminae indet. Bonaparte 1853 seriema
MPM-PV 3512, tibiotarsus fragment