Keeline ( of the United States)

Also known as Quarry Z

Where: Niobrara County, Wyoming (43.0° N, 104.5° W: paleocoordinates 43.9° N, 99.4° W)

• coordinate based on political unit

• small collection-level geographic resolution

When: Harrison Formation, Harrisonian (24.8 - 20.4 Ma)

• said to be late Arikareean

Environment/lithology: terrestrial

Primary reference: J. H. Wahlert and R. A. Souza. 1988. Skull morphology of Gregorymys and relationships of the Entoptychinae (Rodentia, Geomyidae). American Museum Novitates 2922:1-13 [J. Alroy/J. Alroy/J. Marcot]more details

PaleoDB collection 17794: authorized by John Alroy, entered by John Alroy on 07.06.1994, edited by Jonathan Marcot

Creative Commons license: CC BY (attribution)

Taxonomic list

• on the Pine Ridge Escarpment according to Schultz and Falkenbach 1950
virtually all Keeline records appear to be from a single F:AM "North of Keeline" locality
Mammalia
 Rodentia - Geomyidae
Gregorymys formosus Matthew 1907 pocket gopher
 Perissodactyla - Rhinocerotidae
Diceratherium armatum2 Marsh 1875 rhinoceros
"north of Keeline, Niobrara County, Wyoming"
Diceratherium annectens10 Marsh 1873 rhinoceros
 Perissodactyla - Tapiridae
Nexuotapirus marslandensis1 Schoch 1984 tapir
"Lusk area, north of Keeline, Niobrara County"
 Perissodactyla - Equidae
Kalobatippus avus8 Schlaikjer 1935 anchitheriine horse
 Carnivora - Amphicyonidae
Mammacyon ferocior n. sp.6 Hunt 2011 bear-dog
"Arikaree Formation, N of Keeline"
 Carnivora - Mustelidae
Megalictis ferox7 Matthew 1907 mustelid carnivore
"North of Keeline (Quarry Z)... Harrison"
Zodiolestes sp.7 Riggs 1942 mustelid carnivore
 Carnivora - Canidae
Enhydrocyon crassidens14 Matthew 1907 canid
North of Keeline
Paraenhydrocyon josephi14 Cope 1881 canid
North of Keeline
Paraenhydrocyon wallovianus14 Cope 1881 canid
North of Keeline, "Lower or Upper Arikaree Group"
Leptocyon gregorii13 Matthew 1907 canine
north of Keeline, middle brown sand
Cormocyon haydeni15 Wang et al. 1999 bone-crushing dog
"north of Keeline"
 Ungulata - Merycoidodontidae
Desmatochoerus megalodon3, "Merycochoerus carrikeri" = Promerycochoerus carrikeri3, "Merycochoerus chelydra" = Promerycochoerus chelydra3, Phenacocoelus typus12, "Merycoides harrisonensis" = Paramerychyus harrisonensis3, "Merychyus crabilli" = Merychyus calaminthus11
Desmatochoerus megalodon3 Peterson 1907 oreodont
"Merycochoerus carrikeri" = Promerycochoerus carrikeri3 Peterson 1907 oreodont
"Merycochoerus chelydra" = Promerycochoerus chelydra3 Cope 1884 oreodont
Phenacocoelus typus12 Peterson 1907 oreodont
Harrison Fm. N of Keeline
"Merycoides harrisonensis" = Paramerychyus harrisonensis3 Peterson 1907 oreodont
"Merychyus crabilli" = Merychyus calaminthus11 Jahns 1940 oreodont
"N. of Keeline" in Niobrara Co.
 Ungulata - Camelidae
Stenomylus keelinensis5 Frick and Taylor 1968 camel
N of Keeline; S. gracilis of Frick 1937
 Ungulata - Protoceratidae
Protoceras neatodelpha n. sp.9 Patton and Taylor 1973 protoceratid
N of Keeline, Niobrara Co., in Harrison Fm. "equivalent"
 Ungulata - Hypertragulidae
Nanotragulus ordinatus4 Matthew 1907 ruminant
"Keeline, Wyoming"