Burmese amber, Kachin amber (EMTG collection) (Cretaceous of Myanmar)

Where: Myanmar (26.4° N, 96.7° E: paleocoordinates 12.4° N, 93.8° E)

• coordinate stated in text

When: Early/Lower Cenomanian (99.6 - 93.5 Ma)

Environment/lithology: terrestrial; amber

Size classes: macrofossils, mesofossils

Preservation: original chitin, amber

Collection methods: Repository: Eventually in the established Three Gorges Entomological Museum, Chongqing, China. Specimens currently on long-term loan in the Institute of Zoology, Chinese Academy of Sciences (IZAS) or to China Agricultural University (CAU).

Primary reference: S. Boucher, M. Bai, B. Wang, W. W. Zhang, and X. K. Yang. 2016. †Passalopalpidae, a new family from the Cretaceous Burmese amber, as the possible sister group of Passalidae Leach (Coleoptera: Scarabaeoidea). Cretaceous Research 64:67-78 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 178121: authorized by Matthew Clapham, entered by Matthew Clapham on 19.04.2016

Creative Commons license: CC BY (attribution)

Taxonomic list

 Euonychophora - Peripatidae
Cretoperipatus burmiticus28 Engel and Grimaldi 2002
EMTG BU001467, BU-001468
Insecta
 Alienoptera - Alienopteridae
Caputoraptor elegans2 Bai et al. 2018 winged insect
EMTG BU-002371, 001054, 002301, 002366
Alienopterus brachyelytrus n. gen. n. sp.1 Bai et al. 2016 winged insect
EMTG BU-001057
 Phasmatodea - Pterophasmatidae
Meniscophasma erythrosticta n. gen. n. sp.37, Pterophasma erromera n. gen. n. sp.37
Meniscophasma erythrosticta n. gen. n. sp.37 Yang et al. 2019 stick insect
EMTG BU-001967
Pterophasma erromera n. gen. n. sp.37 Yang et al. 2019 stick insect
EMTG BU-001438
 Phasmatodea - Archipseudophasmatidae
Pseudoperla scapiforma6 Chen et al. 2017 stick insect
EMTG BU-001959
Pseudoperla leptoclada n. sp.7 Chen et al. 2017 stick insect
EMTG BU-001003
 Phasmatodea - Timematidae
Tumefactipes prolongates n. gen. n. sp.6
Tumefactipes prolongates n. gen. n. sp.6 Chen et al. 2019 stick insect
EMTG BU-001232
 Embioptera - Clothodidae
Gnethoda odontophora n. sp.9 Lai et al. 2021 webspinners
EMTG BU-001
 Embioptera - Scelembiidae
Multivena curvivena n. gen. n. sp.9
Multivena curvivena n. gen. n. sp.9 Lai et al. 2021 webspinners
EMTG BU-023
 Zoraptera - Zorotypidae
"Zorotypus oligophleps n. sp." = Zorotypus (Zorotypus) denticulatus20, "Zorotypus robustus n. sp." = Zorotypus (Octozoros) cenomanianus20
"Zorotypus oligophleps n. sp." = Zorotypus (Zorotypus) denticulatus20 Yin et al. 2017 winged insect
EMTG BU-002294
"Zorotypus robustus n. sp." = Zorotypus (Octozoros) cenomanianus20 Yin et al. 2017 winged insect
EMTG BU-002293
 Grylloblattodea -
Sylvalitoralis cheni n. sp.41 Zhang et al. 2016 ice crawler
EMTG BU-001678
 Dicondylia - Chresmodidae
Chresmoda chikuni n. sp.40 Zhang and Ge 2017 winged insect
EMTG BU-002027, 001010
 Permopsocida - Archipsyllidae
Burmopsylla maculata n. gen. n. sp.12
Burmopsylla maculata n. gen. n. sp.12 Liang et al. 2016 winged insect
EMTG BU-001053, 001056, 001087
 Hemiptera - Tettigarctidae
Burmaprosbole nervosa n. gen. n. sp.30
Burmaprosbole nervosa n. gen. n. sp.30 Qiao et al. 2021 hairy cicada
EMTG BU-00002069
 Psocodea - Archimenoponidae
Archimenopon myanmarensis n. gen. n. sp.42
Archimenopon myanmarensis n. gen. n. sp.42 Zhang et al. 2024 parasitic louse
EMTG BU-MA002385-1, 2
 Mecoptera - Bittacidae
Bittacus lepiduscretaceus n. sp.11 Li et al. 2018 hangingfly
EMTG BU-001263
 Raphidioptera - Mesoraphidiidae
Dolichoraphidia aspoecki n. gen. n. sp.15 Liu et al. 2016 snakefly
EMTG BA-001002, 002179, 002253
Dolichoraphidia engeli n. sp.15 Liu et al. 2016 snakefly
EMTG BA001300, 001252
Nanoraphidia electroburmica15 Engel 2002 snakefly
EMTG BA-001202, 002164
Rhynchoraphidia burmana n. gen. n. sp.15 Liu et al. 2016 snakefly
EMTG BA-002256
Burmoraphidia reni n. gen. n. sp.15 Liu et al. 2016 snakefly
EMTG BA-001272, 001772, 002135, 002158, 002197, 002260, 002254, 002255
 Raphidioptera - Baissopteridae
Baissoptera burmana23 Lu et al. 2020 snakefly
EMTG BU-002258, 002190, 002104
 Neuroptera - Dipteromantispidae
Halteriomantispa grimaldii n. gen. n. sp.16, Mantispidipterella longissima n. gen. n. sp.17
Halteriomantispa grimaldii n. gen. n. sp.16 Liu et al. 2016 lacewing
EMTG BU-001924
Mantispidipterella longissima n. gen. n. sp.17 Liu et al. 2016 lacewing
EMTG BU-002370
 Neuroptera - Berothidae
Cornoberotha anomala n. gen. n. sp.36 Yang et al. 2019 beaded lacewing
EMTG BU001249
Cornoberotha aspoeckae36 Yang et al. 2019 beaded lacewing
EMTG BU001214
Dolichoberotha bifurcata36 Yang et al. 2019 beaded lacewing
EMTG BU002117, BU001991
Xiaoberotha bipunctata n. gen. n. sp.34 Shi et al. 2019 beaded lacewing
EMTG BU-001590
 Neuroptera - Mantispidae
Doratomantispa pubescens n. sp.22 Lu et al. 2020 mantidfly
EMTG BU-001759
Acanthomantispa immaculata n. gen. n. sp.22 Lu et al. 2020 mantidfly
EMTG BU-001408
Acanthomantispa maculata n. sp.22 Lu et al. 2020 mantidfly
EMTG BU-002131, 002136, 002270
Parasymphrasites electrinus n. gen. n. sp.22 Lu et al. 2020 mantidfly
EMTG BA-002162
 Neuroptera - Corydasialidae
Megalopteroneura chenxingi n. gen. n. sp.18
Megalopteroneura chenxingi n. gen. n. sp.18 Liu et al. 2016 lacewing
EMTG BU-001023
 Neuroptera - Mesochrysopidae
Pedanoptera arachnophila n. gen. n. sp.19
Pedanoptera arachnophila n. gen. n. sp.19 Liu et al. 2016 lacewing
EMTG BU-001049, BU001066; larvae BU-001111, 001113
 Neuroptera - Babinskaiidae
Burmobabinskaia tenuis n. gen. n. sp.27, Electrobabinskaia burmana n. gen. n. sp.27
Burmobabinskaia tenuis n. gen. n. sp.27 Lu et al. 2016 lacewing
EMTG BU-002356
Electrobabinskaia burmana n. gen. n. sp.27 Lu et al. 2016 lacewing
EMTG BU-001698
 Neuroptera - Myrmeleontoidae
Allopteroneura burmana n. gen. n. sp.21
Allopteroneura burmana n. gen. n. sp.21 Lu et al. 2019 lacewing
EMTG BU-002082
 Neuroptera -
Fiaponeura penghiani n. gen. n. sp.26
Fiaponeura penghiani n. gen. n. sp.26 Lu et al. 2016 lacewing
EMTG BU-001679
 Neuroptera - Kalligrammatidae
Cretanallachius magnificus14 Huang et al. 2015 lacewing
EMTG BU-002169, 002266
Burmopsychops limoae n. gen. n. sp.26 Lu et al. 2016 lacewing
EMTG BU-001293
 Neuroptera - Psychopsidae
Electropsychops handlirschi n. gen. n. sp.25
Electropsychops handlirschi n. gen. n. sp.25 Lu et al. 2017 silky lacewing
EMTG BU-001914
 Neuroptera - Ithonidae
Burmithone pennyi n. gen. n. sp.24
Burmithone pennyi n. gen. n. sp.24 Lu et al. 2017 moth lacewing
EMTG BU-002284
 Neuroptera - Dilaridae
Dilar cretaceus n. sp.13, Cretodilar burmanus n. gen. n. sp.13
Dilar cretaceus n. sp.13 Liu and Zhang 2016 pleasing lacewing
EMTG BU-001748
Cretodilar burmanus n. gen. n. sp.13 Liu and Zhang 2016 pleasing lacewing
EMTG BU-001752
 Neuroptera - Coniopterygidae
Achlyoconis heptatrichia10 Engel 2016 dustywing
EMTG BU-001474, 001934, 01199
 Coleoptera - Ripiphoridae
Ripiphoridae indet.29 Gemminger 1870 wedge shaped beetle
EMTG BU-001009
Ripidiinae indet.5 Gerstaecker 1855 wedge shaped beetle
EMTG BU-002386
 Coleoptera - Endomychidae
Cretaparamecus tarsalis n. gen. n. sp.35
Cretaparamecus tarsalis n. gen. n. sp.35 Tomaszewska et al. 2018 handsome fungus beetle
EMTG BU-001534
 Coleoptera - Cantharidae
Sanaungulus emarginaticollis n. sp.38, "Burmomiles oblongoculus n. sp." = Elektrokleinia oblongoculus39
Sanaungulus emarginaticollis n. sp.38 Yang et al. 2022 soldier beetle
EMTG BU001881
"Burmomiles oblongoculus n. sp." = Elektrokleinia oblongoculus39 Yang et al. 2021 soldier beetle
EMTG BU001426
 Coleoptera - Staphylinidae
Clidicostigus arachnipes n. gen. n. sp.8
Clidicostigus arachnipes n. gen. n. sp.8 Jałoszyński et al. 2017 antlike stone beetle
EMTG BU-001212
 Coleoptera - Passalopalpidae
Passalopalpus cheni Boucher et al. 2016 scarab beetle
BU-001782, 001897, 002077, 002119, 002207
 Coleoptera - Lucanidae
Electraesalopsis beuteli n. gen. n. sp.31
Electraesalopsis beuteli n. gen. n. sp.31 Bai et al. 2017 stag beetle
EMTG BU-001838
 Coleoptera - Hybosoridae
Hybosorus ocampoi n. sp.4 Bai and Zhang 2016 scavenger scarab beetle
EMTG BU-001785
 Coleoptera - Bolboceratidae
Amberathyreus beuteli n. gen. n. sp.3
Amberathyreus beuteli n. gen. n. sp.3 Bai and Zhang 2017 dor beetle
EMTG BU-002290
 Dicondylia -
Strepsiptera indet.29 Kirby 1813 twisted-winged parasite
EMTG BU-002386
Arachnida
 Opiliones - Epedanidae
Petrobunoides sharmai n. gen. n. sp.32
Petrobunoides sharmai n. gen. n. sp.32 Selden et al. 2016 harvestman
EMTG BU-002036
 Araneae - Tetrablemmidae
Electroblemma bifida n. gen. n. sp.33
Electroblemma bifida n. gen. n. sp.33 Selden et al. 2016 armored spider
EMTG BU-001055, 001725