Golyusherma, PIN 4276 (Permian to of Russian Federation)

Where: Udmurtia, Russian Federation (56.0° N, 52.7° E: paleocoordinates 28.2° N, 44.2° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Parabradysaurus silantjevi other zone, Baytugan Formation, Ufimian to Ufimian (279.5 - 268.0 Ma)

• Golyusherma Subassemblage of the Ocher Assemblage; terminal part of the Ufimian to the early part of the Late Kazanian (Golubev 2000).

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; gray sandstone

Size class: macrofossils

Collected by Golubev in 1990; reposited in the PIN

Primary reference: V. K. Golubev. 1995. New species of Melosaurus (Amphibia, Labyrinthodontia) from the Kazanian of the Kama River drainage area. Paleontological Journal 29(3):107-119 [R. Butler/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 187322: authorized by Richard Butler, entered by Richard Butler on 20.07.2017, edited by Joseph Flannery-Sutherland

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
unclassified
  -
Striatohaplopinites perfectus1
Limitisporites
  -
Limitisporites sp.1
Limitisporites sp.1 Leschik 1956
Sphenopsida
  -
Paracalamites sp.1
Paracalamites sp.1 Zalessky 1927
Osteichthyes
 Elonichthyiformes - Acrolepidae
Acropholis sp.1, Acropholis kamensis1, Acropholis stensioei1, Plegmolepis sp.2, Watsonichthys sp.2, Palaeostrugia rhombifera2
Acropholis sp.1 Aldinger 1937 ray-finned fish
Acropholis kamensis1 Esin 1995 ray-finned fish
Acropholis stensioei1 Aldinger 1937 ray-finned fish
Plegmolepis sp.2 Aldinger 1937 ray-finned fish
Watsonichthys sp.2 Aldinger 1937 ray-finned fish
Palaeostrugia rhombifera2 Eichwald 1860 ray-finned fish
 Elonichthyiformes - Elonichthyidae
Elonichthys contortus2
Elonichthys contortus2 Esin 1995 ray-finned fish
 Elonichthyiformes - Acrolepididae
Kazanichthys golyushermensis1, Acrolepis sp.2, "Acrolepis cf. sedgwicki" = Acrolepis sedgwickii2
Kazanichthys golyushermensis1 Esin 1995 ray-finned fish
Acrolepis sp.2 Agassiz 1833 ray-finned fish
"Acrolepis cf. sedgwicki" = Acrolepis sedgwickii2 Agassiz 1833 ray-finned fish
Actinopteri
 Semionotiformes - Semionotidae
Acentrophorus sp.2, Acentrophorus varians2
Acentrophorus sp.2 Traquair 1877
Acentrophorus varians2 Kirkby 1862
Osteichthyes
 Palaeonisciformes - Boreolepidae
Boreolepis jenseni2
Boreolepis jenseni2 Aldinger 1937
 Palaeonisciformes - Palaeoniscidae
Grygorichthys sp.2, Koinichthys ivachenkoi2, Palaeoniscum kazanense2
Grygorichthys sp.2 Esin 1990
Koinichthys ivachenkoi2 Nurgaliev et al. 2015
 Palaeonisciformes - Pygopteridae
cf. Pygopterus sp.2
cf. Pygopterus sp.2 Agassiz 1833
 Palaeonisciformes - Platysomidae
Platysomus sp.2, Platysomus cf. striatus2, Eurysomus sp.2, cf. Kargalichthys sp.2, "Wardichthys inobilis" = Kargalichthys pritokensis2
Platysomus sp.2 Agassiz 1833
Platysomus cf. striatus2 Agassiz 1833
Eurysomus sp.2 Young 1866
cf. Kargalichthys sp.2 Minikh 1986
"Wardichthys inobilis" = Kargalichthys pritokensis2 Minikh 1992
 Paramblypteriformes -
Paramblypterus sp.2
Paramblypterus sp.2 Blot 1966
Amphibia
 Temnospondyli - Archegosauridae
Platyoposaurus sp., Melosaurus compilatus n. sp.
Platyoposaurus sp. Lydekker 1889 tetrapod
PIN No. 4276/37
Melosaurus compilatus n. sp. Golubev 1995 tetrapod
PIN No. 4276/60 (holotype), fragment of skull roof. Nos. 4276/1, 18, 51, 73 - angularia; 2, 5, 10, 26, 39, 40, 56 - claviculae; 61 - coronoideum; 8, 11, 12, 36, 41, 48 - costae; 23, 24, 29, 47, 55, 65, 70 - dentalia; 22 - ectopterygoideum; 33, 43, 50 - frontalia; 46 - ilium; 16 - interclavicula; 31, 66, 69 - jugalia; 25, 38, 57 - maxillaria; 6 - nasale; 63 - palatinum; 54,, 67 - parietalia; 34 - postorbitale; 19 - postparietale; 14, 21, 62 - postsplenialia; 4 - praearticulare; 17, 28, 53 - praefrontalia; 30, 71, 72 - praemaxillaria; 20, 45 - pterygoidea; 68 - quadratojugale; 15 - quadratum; 44, 59 - scapulocoracoidea; 35 - spleniale; 64 - tabulare; 9, 58 - vomeres.
Osteichthyes
 Therapsida - Anteosauridae
Microsyodon orlovi n. gen. n. sp.
Microsyodon orlovi n. gen. n. sp. Ivakhnenko 1995 therapsid
PIN No. 4276/13
 Seymouriamorpha - Kotlassidae
cf. Leptoropha talonophora
cf. Leptoropha talonophora Chudinov 1955 tetrapod
PIN No. 4276/27
Bivalvia
  -
Bivalvia indet.1
Bivalvia indet.1 Linnaeus 1758 clam
1Bakaev 2022; 2Golubev 2001