Lipovskaya gully (Donskaya Luka) (Triassic of Russian Federation)

Also known as Lipovaya Balka

Where: Volgograd, Russian Federation (49.3° N, 43.7° E: paleocoordinates 25.8° N, 40.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Lipovskaya Formation, Olenekian (251.2 - 247.2 Ma)

• Lower Triassic, equivalent of Yarengian Gorizont, Lipovskaya Svita (Novikov, 1991, Paleont. Zh. 1991(2)).

•Lipovskaya Svita, Gamian Gorizont, Yarengian Supergorizont, Upper Olenekian Substage, Olenekian Stage, Lower Triassic (Arkhangelskii & Sennikov, 2008).

•The Yarengian is uppermost Olenekian according to fig. 2 in Tverdokhlebov et al. (2003, Earth Sci. Rev. 60).

•Chronostratigraphy originally entered as >>Early/Lower Triassic<< is rendered more precisely according to data given above.

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Reposited in the PIN

Primary reference: M. F. Ivakhnenko. 1973. New Cisuralian cotylosaurs. Paleontological Journal 7(2):247-249 [J. Alroy/R. Whatley/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 27343: authorized by John Alroy, entered by Robin Whatley on 02.12.2002, edited by Torsten Liebrecht, Richard Butler and Jonathan Tennant

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Chondrichthyes
 Hybodontiformes - Hybodontidae
Hybodus maximi4 elasmobranch
 Hybodontiformes - Lonchidiidae
Lissodus triaktis4 elasmobranch
Lissodus angulatus4 Stensiö 1921 elasmobranch
Actinopteri
 Saurichthyiformes - Saurichthyidae
Saurichthys sp.4 Agassiz 1843
 Parasemionotiformes - Parasemionotidae
Watsonulus sp.4 Brough 1939
Osteichthyes
 Dipnoi - Ceratodontidae
Ceratodus jechartiensis4 Minikh 1977 lungfish
? Microceratodus sp.4 Texeira 1954 lungfish
 Dipnoi - Ptychoceratodontidae
"Ceratodus donensis" = Ptychoceratodus donensis4
"Ceratodus donensis" = Ptychoceratodus donensis4 Vorobyeva and Minikh 1968 lungfish
 Dipnoi - Arganodontidae
"Ceratodus multicristatus" = Arganodus multicristatus4
"Ceratodus multicristatus" = Arganodus multicristatus4 Vorobyeva and Minikh 1968 lungfish
subspecies: Ceratodus multicristatus lipovensis
Coelacanthimorpha
 Coelacanthiformes - Latimeriidae
? Holophagus sp.4 Egerton 1861 coelacanth
Osteichthyes
 Temnospondyli - Trematosauridae
Trematosaurus galae n. sp.4 Novikov 2010 tetrapod
PIN 1043/774, preorbital skull fragment without the narial region (holotype). Eight fragments of the postorbital skull region (specimens PIN, nos. 1043/59, 75, 779, 783, 867, 976, 996, 1089); four premaxillae (PIN, nos. 1043/54, 104, 105, 810); one maxillary fragment (PIN, no. 1043/370); one postfrontal (PIN, no. 1043/977); two postorbitals (PIN, nos. 1043/886, 978); five complete and fragmentary jugals (PIN, nos. 1043/880, 912, 1050, 1093, 1097); 11 complete and fragmentary squamosals (PIN, nos. 1043/124, 342, 343, 950, 998, 999, 1084–1088); five complete and fragmentary quadratojugals (PIN, nos. 1043/347, 351, 357, 972, 973); four postparietals (PIN, nos. 1043/195, 869, 951, 1091); 27 complete and fragmentary tabulars (PIN, nos. 1043/52, 92, 94, 95, 96, 115–120, 303, 758, 793, 806–808, 903, 904, 952–955, 957–959, 997); five vomer fragments (PIN, nos. 1043/674, 891, 892, 1004, 1098); six complete and fragmentary palatines (PIN, nos. 1043/123, 775, 877, 878, 915, 1099); nine skull base fragments (PIN, nos. 1043/77, 655, 672, 673, 875, 893, 987, 988, 990); eight complete and fragmentary pterygoids (PIN, nos. 1043/101, 121, 792, 923, 949, 991–993); eight complete and fragmentary parasphenoids (specimen PIN, nos. 1043/107, 109, 390, 675, 872–874, 906); nine complete and fragmentary exoccipitals (PIN, nos. 1043/69, 70, 76, 93, 97, 98, 127, 501, 989); and a fragmentary right stapes (PIN, no. 1043/678).
 Temnospondyli -
 Temnospondyli - Rhytidosteidae
Rhytidosteidae indet.4 Huene 1920 tetrapod
 Temnospondyli - Tupilakosauridae
Batrachosuchoides sp.9 Shishkin 1967 tetrapod
 Tetrapoda - Bystrowianidae
Dromotectum abditum n. sp.10 Shishkin et al. 2014 tetrapod
Reptilia
 Procolophonia - Procolophonidae
? Kapes sp.4, "Tichvinskia enigmatica n. sp." = Orenburgia enigmaticus2
? Kapes sp.4 Ivakhnenko 1975 parareptile
"Tichvinskia enigmatica n. sp." = Orenburgia enigmaticus2 Chudinov and Vjushkov 1956 parareptile
PIN 1043/1 (type)
 Eosuchia -
Coelodontognathus donensis n. gen. n. sp.1 Ochev 1967 archosauromorph
PIN 4173/129 (SGU 104/3103) (type), PIN 4173/130 (SGU 104/3104); both are dentaries
Coelodontognathus ricovi n. sp.1 Ochev 1967 archosauromorph
PIN 4173/127 (SGU 104/3101) (type), PIN 4173/128 (SGU 104/3102); both are dentaries
Tsylmosuchus donensis n. sp.5 Sennikov 1990 archosauromorph
PIN 1043/42, cervical vertebra (holotype)
 Loricata - Rauisuchidae
Scythosuchus basileus n. gen. n. sp.5
Scythosuchus basileus n. gen. n. sp.5 Sennikov 1999 rauisuchid
PIN 1043/144, fragmentary cervical vertebra (holotype). PIN 1043/537, 538, 540, 607, and 608, fragmentary postorbitals; PIN 1043/574 and 615–617, fragmentary maxillae; PIN 1043/142 and 612, left squamosals; PIN 1043/525, articular part of the left ramus of the lower jaw; PIN 1043/575–584 and 636–639, teeth; PIN 1043/393, spinous process of the epistropheus; PIN 1043/143 and 630, spinous processes of the cervical vertebrae; PIN 1043/506 and 507, spinous processes of the posterior cervical vertebrae; PIN 1043/656, cervical vertebra; PIN 1043/657 and 658, posterior cervical and anterior thoracic vertebrae; PIN 1043/532, 609, and 659, spinous processes of anterior thoracic vertebrae; PIN 1043/533, 621, and 622, spinous processes of posterior thoracic vertebrae; PIN 1043/146–176 and 531, thoracic vertebrae; PIN 1043/660, sacral vertebra; PIN 1043/548–573 and 631–634, caudal vertebrae; PIN 1043/511, proximal fragment of the left humerus; PIN 1043/652, fragmentary right ilium; PIN 1043/517, 518, and 625 fragmentary femurs; PIN 1043/515 right tibia; PIN 1043/516, fragmentary fibula; PIN 1043/547 right calcaneum; PIN 1043/544–546 distal tarsal elements.
 Pseudosuchia - Ctenosauriscidae
Bystrowisuchus flerovi n. gen. n. sp.8
Bystrowisuchus flerovi n. gen. n. sp.8 Sennikov 2012 archosaur
PIN 1043/1346, anterior thoracic vertebra (holotype). PIN 1043/393, neural spine of epistropheus; PIN 1043/1128, incomplete cervical vertebra; PIN 1043/ 656, 1493, incomplete posterior cervical vertebrae; PIN 1043/147, incomplete anterior thoracic vertebra; PIN 1043/831, right ilium
 Eosuchia - Tanystropheidae
Augustaburiania vatagini n. gen. n. sp.7
Augustaburiania vatagini n. gen. n. sp.7 Sennikov 2011 archosauromorph
PIN 1043/587, cervical vertebra (holotype). PIN no. 1043/842, epistropheus; PIN, nos. 1043/840, 841, 1141, 1327, 1328, 1392, cervical vertebrae; PIN, nos. 1043/588, 651, fragments of cervical vertebrae; PIN, nos. 1043/585, 586, 843, 1142, posterior cervical vertebrae; PIN, nos. 1043/589–594, 844, 845, 1143, 1144, 1310, pectoral vertebrae; PIN, nos. 1043/397, 398, 598, neural arches of pectoral vertebrae; PIN, no. 1043/595, sacral vertebra; PIN, nos. 1043/596, 597, 847–850, 1145–1147, caudal vertebrae; PIN, no. 1043/648, fragment of right humerus; PIN, no. 1043/771, fragment of left humerus; PIN, nos. 1043/649, 650, 851, left femora; PIN, nos. 1043/603, 671, proximal fragments of left femora; PIN, no. 1043/606, proximal fragment of right femur; PIN, nos. 1043/601, 604, 641, 642, distal fragments of femora; PIN, no. 1043/514, articulated distal fragment of femur and proximal fragment of tibia (left); PIN, nos. 1043/512, 513, 644–646, 770, proximal fragments of tibiae; PIN, no. 1043/643, distal fragment of tibia; PIN, no. 1043/647, proximal fragment of fibla; and PIN, nos. 1043/602, 1393, metatarsals.
 Sauropterygia - Cymatosauridae
Tanaisosaurus kalandadzei n. gen. n. sp.6
Tanaisosaurus kalandadzei n. gen. n. sp.6 Sennikov 2001 diapsid
Osteichthyes
  -
Vitalia grata n. gen. n. sp.
Vitalia grata n. gen. n. sp. Ivakhnenko 1973 amniote
PIN 4173/126 (SGU 104/3105) (type; prevoiusly included in the hypodigm of Coelodontognathus donensis by Ochev, 1967), PIN 1043/627, 1043/628; all are dentaries; current catalogue number of type and catalogue numbers of referred specimens are from Arkhangelskii & Sennikov (2008)
Mammalia
 Trogontia - Esthonychidae
Putillosaurus sennikovi n. gen. n. sp.3
Putillosaurus sennikovi n. gen. n. sp.3 Surkov 2005 whale
PIN 1043/614 (holotype)
 Polyprotodonta - Herpetotheriidae
Doniceps lipovensis5 Otschev and Rykov 1968 marsupial
Saratov University 104/3106, premaxilla (specimen lost)