Also known as Locality 34 (Romer 1928), faunal Zone 4, Lower Clear Fork Group, Texas red-beds, redbeds; Locality VIIa (Romer & Price 1940); "Labidosaurus pocket"; Coffee Creek Bonebed; West Coffee Creek; Middle Coffee Creek; East Coffee Creek
Where: Baylor/Willbarger County, Texas (33.8° N, 99.2° W: paleocoordinates 2.3° N, 28.7° W)
• coordinate based on nearby landmark
• local area-level geographic resolution
When: Arroyo Formation (Clear Fork Group), Kungurian (279.3 - 272.3 Ma)
• "probably not much over 100 feet above the Lueders limestone; the lowest portion may have lain below it.." Details for Faunal Zone 4 of Romer 1928: "The vertebrate remains are confined to approximately the lowest 250 feet or so of the Clear Fork.."
•the base and top of the Clear Fork Group (i.e., Arroyo, Vale, and Choza formations) are both within the Kungurian: see Wardlaw 2005 (Permophiles) for the base, and DiMichele et al. 2001 (J Paleont) for the top
• group of beds-level stratigraphic resolution
Environment/lithology: terrestrial; red claystone and conglomeratic sandstone
Size class: macrofossils
Collected by J. W. Kitching, P. C. Miller, E. C. Olson, C. H. Sternberg, S. W. Williston, among others in 1895, 1897, 1909, 1965, 1972; reposited in the AMNH, FMNH, MCZ, USNM
• Romer (1928) paper is a review of vertebrates from the Texas red-beds with previously unreported (or undetailed) localities.
•Additional institutions storing material from Coffee Creek:
•BP (Bernard Price Institute, University of the Witwatersrand, Johannesburg, South Africa)
•BSP (Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany)
•UMMP (University of Michigan Museum of Paleontology, Ann Arbor, MI)
Primary reference: E. D. Cope. 1882. Third contribution to the history of the Vertebrata of the Permian formation of Texas. Proceedings of the American Philosophical Society 20:447-461 [M. Carrano/M. Carrano/M. Carrano]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 28256: authorized by John Alroy, entered by Robin Whatley on 03.02.2003, edited by Torsten Liebrecht, Matthew Carrano, Emma Dunne and Richard Butler
Creative Commons license: CC BY (attribution)
Taxonomic list
Amphibia | |
"Isodectes obtusus" = Isodectes obtusus19 Cope 1868 tetrapod AMNH 4744 (Originally A. ? casei figured by Broili 1913, pl. 9, figs 1-2, presumed destroyed in 1944)
"? Acheloma casei n. sp." = Isodectes obtusus2 Cope 1868 tetrapod BSP unknown catalogue number; occurence is deducted from the original 2 specimens of Acheloma sp. reported in the primary reference
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Trimerorhachis mesops n. sp.17
Trimerorhachis mesops n. sp.17 Cope 1896 tetrapod Authority is not Romer 1928. entered as "Trimerorachis"
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Dissorophus multicinctus11, Aspidosaurus chiton n. gen. n. sp.17, "Alegeinosaurus aphthitos n. gen. n. sp." = Aspidosaurus aphthitos19
"Alegeinosaurus aphthitos n. gen. n. sp." = Aspidosaurus aphthitos19 Cope 1881 tetrapod AMNH 4756 (holotype)
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"Anisodexis imbricarius n. gen. n. sp." = Eryops megacephalus Cope 1877 tetrapod | |
cf. Zatrachys sp.19 Cope 1878 tetrapod SMU specimens (Murray & Johnson, 1987) from East Coffee Creek
"Zatrachys microphthalmus n. sp." = Dasyceps microphthalmus17 Cope 1869 tetrapod Authority is not Romer 1928. 7 Zatryachys specimens total, see below Z. conchigerus.
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Zatrachys conchigerus n. sp.17
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Diplocaulus sp.15, Diplocaulus brevirostris n. sp.15, "Diplocaulus magnicornis n. sp." = Diplocaulus magnicornis, "Diplocaulus limbatus n. sp." = Diplocaulus magnicornis12
Diplocaulus sp.15 Cope 1877 tetrapod Romer (1928) reports 72 Diplocaulus specimens total; since Olson (1951) refers only 28 specimens from this locality to particular species the remainder (no catalogue numbers given) are listed here as Diplocaulus sp.
"Diplocaulus magnicornis n. sp." = Diplocaulus magnicornis Cope 1882 tetrapod AMNH 4466, 4467, 4469, 4484, 4491, 4494, 4498, 4504, 4511*, 4512*, 4514, 4539 (type); CNHM UC 229, 636, 637, 1650, 1654, 1656-1658, 1660, 1663, P 12689; catalogue numbers are from Olson (1951), asterisks mark specimens with uncertain locality data
"Diplocaulus limbatus n. sp." = Diplocaulus magnicornis12 Cope 1882 tetrapod AMNH 4471 (type); because D. limbatus was found to be an invalid species by Olson (1951) only the type specimen is listed here
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Osteichthyes | |
"Bolbodon tenuitectes n. gen. n. sp." = Diadectes tenuitectes13, "Diadectes huenei n. sp." = Diadectes tenuitectes17
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Dimetrodon loomisi16, "Embolophorus dollovianus n. sp." = Dimetrodon dollovianus10, "Dimetrodon gigashomogenes n. sp." = Dimetrodon giganhomogenes5
Dimetrodon loomisi16 Romer 1937 synapsid FMNH 114 (formerly referred to D. dollovianus) + the rest of the total of 15 Dimetrodon specimens originally reported by Romer (1928)
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"Varanosaurus acutirostris" = Varanosaurus acutirostris18, "Poecilospondylus francisi n. gen. n. sp." = Varanosaurus acutirostris7
"Poecilospondylus francisi n. gen. n. sp." = Varanosaurus acutirostris7 Broili 1904 ophiacodont AMNH 4174 (type)
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Tetraceratops insignis n. gen. n. sp.14
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Reptilia | |
"Pariotichus hamatus n. sp." = Labidosaurus hamatus20, "Labidosaurus hamatus" = Labidosaurus hamatus17, Labidosaurus broilii n. sp.8, "Ectocynodon aguti n. sp." = Captorhinus aguti, "Hypopnous squaliceps n. gen. n. sp." = Captorhinus aguti17, "Pariotichus isolomus n. sp." = Captorhinus aguti12, "Captorhinus angusticeps n. gen. n. sp." = Captorhinus aguti12
"Labidosaurus hamatus" = Labidosaurus hamatus17 Cope 1896 eureptile AMNH 4406, 4414, 4417, 4421, 4427; CNHM ?*, CNHM UR 161, 273, UC ?*, UC 180, 634*, 643, 724-726, 728-730, 732, 1059, 1199, P 12758; CM 73370 (formerly UCLA VP 3167), CM 73371 (formerly UCLA VP 3200), CM 76876; UCLA VP 435, 436, 3115, 3491; USNM 17045; MCZ 8923; catalogue numbers are from Seltin (1959); specimens not reported by Seltin are compiled from Sumida (1987, 1989) and Modesto (2007); asterisks mark specimens of which the locality is not precisely known but come from the Arroyo Fm. of Baylor Co.
"Hypopnous squaliceps n. gen. n. sp." = Captorhinus aguti17 Cope 1882 eureptile AMNH 4335 (type); referred specimen of Captorhinus aguti in Seltin (1959); = "indeterminate romeriid" of Fox & Bowman (1966)
"Captorhinus angusticeps n. gen. n. sp." = Captorhinus aguti12 Cope 1882 eureptile AMNH 4438 (type); 4334, 4340, 4407, 4408, 4410A, 4438, 4443-4445, 4455*, 4457, 4458, 4760, 6790, 6791; CNHM UC 651, 700, 1043, 1702, 1703, UR 274-276, 292, 327, 351, 371, 372, 1206*; MCZ 1059; USNM 17084*; catalogue numbers are from Seltin (1959) and from Fox & Bowman (1966); all are referred specimens of C. aguti in these references and former assignment to C. angusticeps is not confirmed; asterisks mark specimens of which the locality is unknown but come from the Arroyo Fm. of Baylor Co.; at least 5 specimens unnumbered or missing
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Osteichthyes | |
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Quasicaecilia texana n. gen. n. sp.3
Quasicaecilia texana n. gen. n. sp.3 Carroll 1990 amniote Exact location where this specimen was found is unknown. Museum data indicates it was collected by C. H. Sternberg in 1917 from "?Baylor County, Texas; Lower Permian, ?Arroyo Formation". USNM 22079 is the holotype (an isolated skull)
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"Cardiocephalus sternbergi n. gen. n. sp." = Cardiocephalus sternbergi1, "Gymnarthrus willoughbyi n. gen. n. sp." = Cardiocephalus sternbergi6, Pariotichus brachyops17
"Gymnarthrus willoughbyi n. gen. n. sp." = Cardiocephalus sternbergi6 Broili 1904 amniote AMNH 4892 (type)
Pariotichus brachyops17 Cope 1878 amniote AMNH 4760 ("homotype"); type skull of "Captorhinus gregoryi" Broom (see Gregory et al., 1956)
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Pantylus coicodus n. sp.17
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? Micraroter erythrogeios4, Pelodosotis elongatum n. gen. n. sp.4, Ostodolepis brevispinatus n. gen. n. sp.23
Pelodosotis elongatum n. gen. n. sp.4 Carroll and Gaskill 1978 amniote UMMP 11156 (type), formerly assigned to Ostodolepis brevispinatus by Case (1929)
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"Seymouria baylorensis n. gen. n. sp." = Seymouria baylorensis1, "Desmospondylus anomalus n. gen. n. sp." = Seymouria baylorensis22
"Seymouria baylorensis n. gen. n. sp." = Seymouria baylorensis1 Broili 1904 tetrapod AMNH 4342 (type); MCZ 1081, 1083, 1084, 1086; FMNH UR 458; specimen numbers are from Laurin (1996); MCZ 1081, 1083, 1084, 1086 are not counted here because they appear to be already included in collection 22715
"Desmospondylus anomalus n. gen. n. sp." = Seymouria baylorensis22 Broili 1904 tetrapod FMNH UC 664, 665 (both constitute the holotype)
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