Kotelnich, Vjatka River ("Sarmin" series), PIN 2212 (Permian of Russian Federation)

Also known as Kotel'nich; Volki, Volky; Vanyushonki

Where: Kirov, Russian Federation (58.2° N, 48.4° E: paleocoordinates 31.8° N, 40.5° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Deltavjatia vjatkensis other zone, Vanyushonki Member (Urpalov Formation), Early/Lower Severodvinian (265.0 - 252.3 Ma)

• Kotelnich Subassemblage of the Sokolki Assemblage; the early part of the Late Tatarian (early Severodvinian) (Golubev 2000; Benton 2012, Geology).

•"The geological age of the beds of the Kotelnitch locality is younger than that of the zone III (series I, acc. to Kassin --Urdzum series, acc. to A. N. Mazarovitich), and apparently they are synchronous with the lower horizons of Zone IV, or the Pareiasaurian zone, i. e., with the lower layers of the Sarmin series of Mazarovitch." (Efremov, 1940).

•"The detailed geology and stratigraphy of the locality was studied by Coffa (1997 [PhD Thesis]). He recognized the Urpalov Formation at the locality as consisting of four members (Fig. 2), the oldest of which - Vanyushonkov Member (arrow in Fig. 2) composed of red calcareous clays and mudstones yields remains of pareiasaurs. At present, the Vanyushonkov Member correlates to the middle of the Urpalov Formation, Kotel'nich Group, Vishkil'skiy Horizon, Upper Tatarian Stage of the Upper Permian (Coffa 1997). The old Severo-Dvinsky Horizon is no longer in usage due to nomenclature problems." (Kordikova & Khlyupin, 2001).

•Formation name, originally entered as >>Sarmin<<, which never was a lithostratigraphic term, was replaced by >>Urpalov<< (TL).

•Stratigraphic age originally entered as >>Rotliegendes-Zechstein<< was refined: The Kotelnich locality is Lower Severodvinskian according to Ivakhnenko (2008). The Kotelnich Subassemblage of the Sokolki Assemblage and the Deltavjatia vjatkensis Assemblage Zone are regarded as early Upper Severodvinian by Golubev (2005, New Mexico Mus. Nat. Hist. Sci. Bull. 30).

• formation-level stratigraphic resolution

Environment/lithology: terrestrial; brown, red, calcareous claystone and sandstone

• Reddish-brown clay beds, red calcareous clays (Efremov, 1940). This primarily refers to the pareiasaur bearing Vanyushonkov Member of the Urpalov Formation (see Kordikova & Khlyupin, 2001). Olson (1957) and Rubidge & Sidor (2002) also mention sandstone as host rock, probably representing other fossiliferous levels than the Vanyushonkov Member.

Size class: macrofossils

Preservation: mold/impression, concretion

Collected by K. Grekhov, among others in 1935, 1999; reposited in the PIN

• The type material of Pareiasuchus vjatkensis and Anthodon rossicus was collected in 1935. The neonate Deltavjatia reported by Kordikova & Khlyupin (2001) was collected in 1999 by K. Grekhov.

•KPM = Kotel'nich Paleontological Museum (= KMR in Lee, 1997, 2000)

•UMZC = University Museum of Zoology, Cambridge, UK

Primary reference: A. Hartmann-Weinberg. 1937. Pareiasauriden als Leitfossilien [Pareiasaurids as guide fossils]. Problemy paleontologii 213:649-712 [J. Mueller/T. Liebrecht/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 28854: authorized by John Alroy, entered by Robin Whatley on 25.02.2003, edited by Torsten Liebrecht, Marcus Walther, Richard Butler and Bethany Allen

Creative Commons license: CC BY (attribution)

Taxonomic list

• "There are at least another four skulls of this animal [i.e., Deltavjatia vjatkensis]. One is in the Palaeontological Institute, Moscow. The labels have been mixed up, but the catalogue number is probably PIN 2212/3. Another is in the private collection of Mr. Terry Manning. Two more are currently undergoing preparation in the Department of Zoology, Erindale College, University of Toronto (Modesto. 1994)." (Lee, 1997, 2000).
Osteichthyes
 Temnospondyli - Dvinosauridae
Dvinosaurus sp.8 Amalitzky 1921 tetrapod
Reptilia
 Procolophonia - Pareiasauridae
"Anthodon rossicus n. sp." = Deltavjatia rossicus, "Pareiasuchus vjatkensis n. sp." = Deltavjatia rossicus, "Anthodon chlynoviensis n. sp." = Deltavjatia rossicus1
"Anthodon rossicus n. sp." = Deltavjatia rossicus Hartmann-Weinberg 1937 parareptile
PIN 2212/2 (type); catalogue number given in Lee (2000)
"Pareiasuchus vjatkensis n. sp." = Deltavjatia rossicus Hartmann-Weinberg 1937 parareptile
PIN 2212/1 (type); catalogue number given in Lee (1997); taxon referred to as "Pareiasaurus viatkensis" in Olson (1957); PIN 2212/6; UMZC T1321; KPM unnumbered, KPM N11/99 (a neonate, reported by Kordikova & Khlyupin, 2001); for repository of additional material see comments on taxonomic list
"Anthodon chlynoviensis n. sp." = Deltavjatia rossicus1 Hartmann-Weinberg 1937 parareptile
 Parareptilia - Nycteroleteridae
Emeroleter levis n. gen. n. sp.3
Emeroleter levis n. gen. n. sp.3 Ivakhnenko 1997 parareptile
PIN 2212/92, an isolated, dorsoventrally flattened skull. PIN 2212/14, partial skull; PIN 2212/89, fragmentary skull; KPM uncat/E1, complete articulated skeleton; KPM uncat/E2, mostly complete articulated skeleton and skull, part and counter part; KPM uncat/E3, mostly complete skull and fragmentary postcranial remains; KPM uncat/E4, partial articulated postcranial skeleton with partial skull.
 Gorgodontia -
Nochnitsa geminidens n. gen. n. sp.6, Viatkogorgon ivakhnenkoi n. gen. n. sp.4
Nochnitsa geminidens n. gen. n. sp.6 Kammerer and Masyutin 2018 reptile
KPM 310, a nearly complete skull and lower jaws with articulated vertebrae, ribs, and partial right forelimb
Viatkogorgon ivakhnenkoi n. gen. n. sp.4 Tatarinov 1999 reptile
PIN 2212/61, skeleton (type)
Osteichthyes
 Therapsida -
Suminia getmanovi n. gen. n. sp.2
Suminia getmanovi n. gen. n. sp.2 Ivakhnenko 1994 therapsid
PIN 2212/10 (type) - incomplete skeleton, 2212/29-33 - skull; 2212/18, /22, /34-39, /52-54 - jaws; 2212/23-28, /42-51 - individual teeth
 Therocephalia - Ictidosuchidae
Scalopodontes kotelnichi n. gen. n. sp.5
Scalopodontes kotelnichi n. gen. n. sp.5 Tatarinov 2000 therapsid
PIN 2212/16 (holotype), skull
 Therocephalia - Scaloposauridae
Muchia microdenta n. sp.7 Ivakhnenko 2011 therapsid
 Therocephalia - Whaitsiidae
Viatkosuchus sumini n. gen. n. sp.5
Viatkosuchus sumini n. gen. n. sp.5 Tatarinov 1995 therapsid
PIN 2212/13 (holotype), skeleton; PIN 2212/94, maxilla
 Therocephalia - Karenitidae
"Karenites ornamentatus n. gen. n. sp." = Karenites ornamentatus5, "Scalopodon tenuisfrons n. gen. n. sp." = Karenites ornamentatus5
"Karenites ornamentatus n. gen. n. sp." = Karenites ornamentatus5 Tatarinov 1995 therapsid
PIN 2212/60 (holotype), partial skeleton. PIN 2212/20, 21, 40, 59, 93, 96, 98
"Scalopodon tenuisfrons n. gen. n. sp." = Karenites ornamentatus5 Tatarinov 1995 therapsid
PIN 2212/97 (holotype), partial skull
 Therocephalia - Perplexisauridae
"Perplexisaurus foveatus n. gen. n. sp." = Perplexisaurus foveatus5, "Chlynovia serridentatus n. gen. n. sp." = Perplexisaurus foveatus5
"Perplexisaurus foveatus n. gen. n. sp." = Perplexisaurus foveatus5 Tatarinov 1997 therapsid
PIN 2212/15 (holotype), skull; 2212/19, dentary; 2212/55, skull fragment
"Chlynovia serridentatus n. gen. n. sp." = Perplexisaurus foveatus5 Tatarinov 1997 therapsid
PIN 2212/90 (holotype), partial skull
 Therocephalia - Kotelcephalonidae
Kotelcephalon viatkensis n. gen. n. sp.5
Kotelcephalon viatkensis n. gen. n. sp.5 Tatarinov 1999 therapsid
PIN 2212/91 (holotype), skull
 Therocephalia -
Gorynychus masyutinae n. gen. n. sp.7
Gorynychus masyutinae n. gen. n. sp.7 Kammerer and Masyutin 2018 therapsid
KPM 346–349