Where: Basque County, Spain (42.7° N, 2.9° W: paleocoordinates 41.3° N, 6.1° W)
• coordinate based on nearby landmark
When: MP 18 (37.2 - 33.9 Ma)
• biozone MP 18, early middle Ludian (Priabonian)
•European Headonian Land Mammal Age corresponding to the reference level MP 18 (Astibia et al., 2000; Badiola, 2004; Badiola et al., 2009b).
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine; carbonaceous marl and limestone
•The depositional setting was a shallow, low-gradient freshwater lake margin with a peripheral swamp (Iriarte et al., 2003; Badiola et al., 2009a).
•the fossiliferous beds, which consist of palustrine- lacustrine coal-bearing marls and marlstones, are located in one of the oldest lacustrine systems in the Miranda-Treviño Basin (Basque-Cantabrian Region) (Astibia et al., 2000)
Size classes: macrofossils, mesofossils
Primary reference: A. Badiola, H. Astibia, A. Aranburu, X. Pereda Suberbiola, X. Murelaga, C. Sese, M. A. Cuesta, S. Moya-Sola, J. I. Baceta and M. Kohler. 1999. A new continental vertebrate locality from the Upper Eocene of Zambrana (Miranda-Trevino Basin, Alava, Basque Country). IV European Workshop on vertebrate Paleontology, Albarracin, Spain 19 [J. Alroy/E. Leckey/E. Leckey]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 38817: authorized by John Alroy, entered by Erin Leckey on 06.05.2004, edited by Richard Butler and Grace Varnham
Creative Commons license: CC BY (attribution)
Taxonomic list
Amphibia | |
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Mammalia | |
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Microchoerus aff. erinaceus6 Wood 1844 tarsier Right mandible fragment with p3-m3 (MCNA 14518); right mandible fragment with p4-m3 (MCNA 14519).
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Leptolophus "new species"6, Leptolophus sp.2, Leptolophus cuestai n. sp.7, Leptolophus franzeni n. sp.7, "Paranchilophus remyi" = Anchilophus (Paranchilophus) remyi6, Pachynolophus zambranensis n. sp.5, Iberolophus arabensis n. gen. n. sp.4, Palaeotherium "new species"
Leptolophus cuestai n. sp.7 Perales-Gogenola et al. 2021 odd-toed ungulate Holotype: MCNA 9948, right M2; MCNA 9947, right M1; MCNA 9002, incomplete left hemimandible preserving the m1–(m2)–(m3) series; MCNA 9949, incomplete right hemimandible showing the p4– m1–(m2)–(m3) series
Leptolophus franzeni n. sp.7 Perales-Gogenola et al. 2021 odd-toed ungulate Holotype: UPV/EHU-Z4.1092, cranium fragment with right and left C–P2–P3–P4–M1–M2–(M3) series. Fragmented right P2 and P4, and left P2; MCNA 11848, left M3; MCNA 11849, right M3; UPV/EHU- Z4.1041 fragment of right hemimandible with a par- tially erupted m3.
"Paranchilophus remyi" = Anchilophus (Paranchilophus) remyi6 Casanovas and Santafé 1989 odd-toed ungulate fragment of mandible with teeth
Pachynolophus zambranensis n. sp.5 Badiola et al. 2005 odd-toed ungulate MCNA 10657 (holotype). MCNA 10658-10661 (paratypes). 21 referred specimens
Iberolophus arabensis n. gen. n. sp.4 Badiola and Cuesta 2008 odd-toed ungulate MCNA 11953 (holotype). Paratypes: MCNA 10183, skull fragment, showing the right and left M1–M3 molar series; MCNA 11888, left P3–4; MCNA 11906, left P1–2; MCNA 13372, left p4; and MCNA 13373, right p1. Referred material: —MCNA 11904, right P3–4; MCNA 11887, 11893, left P3–4; MCNA 13375, fragment of the left P3–4; MCNA 11905, fragment of the left P3–4; MCNA 11950, right P1-2 (incomplete lingually); MCNA 13374, fragment of the right upper premolar; MCNA 11951, fragment of the upper premolar; MCNA 11952, fragment of the lower premolar
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Reptilia | |
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