Moratines (Miocene of Spain)

Where: Madrid, Spain (40.4° N, 3.7° W: paleocoordinates 40.4° N, 5.2° W)

• coordinate stated in text

When: Orleanian (20.0 - 16.0 Ma)

• Ceratinly older than MN6, probably MN4b, Middle Orleanian, Middle Miocene.

Environment/lithology:

Collection methods: 50 cm by 1m was excavated in a rescue excavation by the 'Equipo de Vertebrados del Instituto de Geologia del CSIC de Madrid.'

Primary reference: M. T. Alberdi, E. Jimenez, J. Morales and C. Sese. 1981. Moratines: pimeros micromamiferos en el Mioceno Medio del area de Madrid. Estudios Geologicos 37:291-305 [A. Turner/H. O'Regan/H. O'Regan]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 40747: authorized by Alan Turner, entered by Hannah O'Regan on 30.06.2004

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Testudines - Testudinidae
"Geochelone bolivari" = Titanochelon bolivari
"Geochelone bolivari" = Titanochelon bolivari Hernandez-Pacheco 1917 turtle
Bones and carapace fragments.
Mammalia
 Carnivora - Percrocutidae
Pseudailurus lorteti carnivoran
1 upper canine.
 Perissodactyla - Equidae
Anchitherium aurelianense Cuvier 1825 anchitheriine horse
2 maxilla fragments, 1 lower M3, 1 lower incisor, plus numerous cranial fragments. 1 distal humerus, 1 astragalus, 3 proximal metapodia, 1 distal metapodial, 2 first phalanges and many carpal and tarsal fragments.
 Perissodactyla - Rhinocerotidae
Rhinocerotidae indet. Gray 1821 rhinoceros
numerous fragments.
 Artiodactyla - Suidae
Suidae indet. Gray 1821 pig
1 mandible with D4-M1 and 1 upper I3.
 Artiodactyla - Palaeomerycidae
Triceromeryx pachecoi Villalta et al. 1946 ruminant
1 maxilla with M1-M3, 1 upper P4, 1 mandible with P4-M1 and 1 distal humerus fragment.
 Artiodactyla - Bovidae
Miotragocerus sp. Stromer 1928 antelope
 Artiodactyla - Cainotheriidae
Cainotherium miocaenicum Crusafont-Pairo 1955 even-toed ungulate
1 upper M3, 1 premolar.
 Lagomorpha - Ochotonidae
Lagopsis penai pika
1 lower P3, 1 lower molar, 1 upper P2, 1 upper molar and 2 molar fragments.
 Rodentia - Cricetidae
Megacricetodon minor Lartet 1851 mouse
1 upper M1. Megacricetodon minor cf. collongensis.
 Rodentia - Gliridae
Armantomys aragonensis de Bruijn 1966 dormouse
Lower teeth: 1 M2, 2 M3. 3 upper M1-2.
Pseudodryomys robustus de Bruijn 1967 dormouse
1 maxilla with P4-M3.
 Rodentia - Sciuridae
Heteroxerus rubricati squirrel
Lower teeth: 2 M1-2, 2 M3. Upper teeth: 1 P4, 3 M1-2, 1 M3.