Talpa (Carboniferous of the United States)

Where: Taos County, New Mexico (36.3° N, 105.6° W: paleocoordinates 1.5° S, 37.5° W)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Flechado Formation, Desmoinesian (312.8 - 306.0 Ma)

• Age estimate of authors is early Desmoinesian based on brachiopods and fusulinids of previous studies. Talpa section, but no beds mentioned.

Environment/lithology: delta front; poorly lithified, fine-grained shale and lithified sandstone

• The Flechado outcrop south of Talpa is predominantly a sequence of steeply dipping, dark gray, fossiliferous marine shale and siltstone units interbedded with thinner beds of conglomeratic to medium-grained, locally cross-bedded, tan to gray sandstones, some bearing plant remains. The lower part of the sequence exposed here, in which fine-grained dark clastics containing abundant fossils predominate (total thickness about three times the cumulative thickness of interbedded sandstones), is about 60 m thick. Above this interval sandstone units become thicker and more numerous, eventually excluding the fossiliferous shale/siltstone beds, and culminating in thick, cliff forming, corss-bedded, channel sand and delta-front bodies.

•JA: fossiliferous facies originally entered as lithified and "silciclastic" but repeatedly referred to as shale by the authors and certainly not well-lithified based on ease of screenwashing the material

Size classes: macrofossils, mesofossils

Reposited in the NMMNH, UNM

Collection methods: bulk, quarrying, surface (in situ), sieve,

• The collection methods were not standardized with some collections made by wet sieving for microgastropods while other horizons were only sampled by surface collecting. The number of specimes are the numbers reported to have been examined for taxonomic purposes. The ecological significance of the number os specimens is unclear.

Primary reference: B. S. Kues and R. L. Batten. 2001. Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology 75(1, supp):1-95 [N. Heim/N. Heim/P. Wagner]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 46536: authorized by Noel Heim, entered by Noel Heim on 18.01.2005, edited by Pete Wagner and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

• 411 juvenile specimens were repored as belonging to the Family Soleniscinae, but not genus identification was made.
Show authors, comments, and common names
Gastropoda
 Cyrtoneritida - Orthonychiidae
"Platyceras (Orthonychia) parvum" = Orthonychia parva
"Platyceras (Orthonychia) parvum" = Orthonychia parva Shumard and Swallow 1858 snail
 Cycloneritida - Trachyspiridae
Trachydomia spp., Trachydomia raymondi, Trachydomia aff. turbonitella, Trachydomia nodosa
Trachydomia spp. Meek and Worthen 1866 snail
juveniles
Trachydomia raymondi Sturgeon 1964 snail
Trachydomia nodosa Meek and Worthen 1860 snail
 Bellerophontida - Bellerophontidae
Pharkidonotus "sp. 1", Pharkidonotus percarinatus, Pharkidonotus aff. megalius3, "Bellerophon crassus" = Waagenella crassus, Bellerophon "sp. 1", Bellerophon spp., Bellerophon cf. incomptus, Bellerophon wewokanus, Patellilabia sp., Retispira tenuilineata, "Knightites montfortianus" = Cymatospira montfortianus, Knightites "sp. 1"
Pharkidonotus "sp. 1" Girty 1912 snail
Pharkidonotus percarinatus Conrad 1842 snail
Pharkidonotus aff. megalius3 Kues 1987 snail
"Bellerophon crassus" = Waagenella crassus Meek and Worthen 1860 snail
Bellerophon "sp. 1" de Montfort 1808 snail
Bellerophon spp. de Montfort 1808 snail
juveniles
Bellerophon cf. incomptus Gurley 1884 snail
Bellerophon wewokanus Girty 1912 snail
Patellilabia sp. Knight 1945 snail
Retispira tenuilineata Gurley 1884 snail
"Knightites montfortianus" = Cymatospira montfortianus Norwood and Pratten 1855 snail
Knightites "sp. 1" Moore 1941 snail
 Bellerophontida - Euphemitidae
Warthia cf. kingi, Euphemites "sp. 1", Euphemites enodis, Euphemites hermosus n. sp., Euphemites aff. kingi, Euphemites nodocarinatus
Warthia cf. kingi Moore 1941 snail
Euphemites "sp. 1" Warthin 1930 snail
Euphemites enodis Sturgeon 1964 snail
Euphemites hermosus n. sp. Kues and Batten 2001 snail
Euphemites aff. kingi Yochelson 1960 snail
Euphemites nodocarinatus Hall 1858 snail
 Heterostropha - Donaldinidae
Donaldina robusta, Donaldina stevensana
Donaldina robusta Stevens 1858 snail
Donaldina stevensana Meek and Worthen 1866 snail
 Heterostropha - Streptacididae
? Streptacis sp., Streptacis whitfieldi
? Streptacis sp. Meek 1871 snail
Streptacis whitfieldi Meek 1871 snail
 Murchisoniina - Murchisoniidae
Bellazona "sp. A"
Bellazona "sp. A" Gordon and Yochelson 1987 snail
 Murchisoniina - Soleniscidae
"Strobeus ? primigenius" = Strobeus primogenius, Soleniscus typicus, "Strobeus regularis" = Soleniscus regularis, Leptoptygma subtilistriatum, Leptoptygma virgatum, Leptoptygma simplex, Strobeus immanis n. sp., Strobeus brevis, Strobeus welleri, Strobeus poromus, Strobeus paludinaeformis, Cylindritopsis vaningeni
"Strobeus ? primigenius" = Strobeus primogenius Conrad 1835 snail
Soleniscus typicus Meek and Worthen 1860 snail
"Strobeus regularis" = Soleniscus regularis Cox 1857 snail
Leptoptygma subtilistriatum Knight 1931 snail
Leptoptygma virgatum Knight 1931 snail
Leptoptygma simplex Knight 1931 snail
Strobeus immanis n. sp. Kues and Batten 2001 snail
Strobeus brevis White 1881 snail
Strobeus welleri Knight 1931 snail
Strobeus poromus Kues 1990 snail
Strobeus paludinaeformis Hall 1858 snail
Cylindritopsis vaningeni Knight 1931 snail
 Murchisoniina - Gosseletinidae
Apachella sp., ? Lunulazona sp., Luciellina ocultabanda n. sp., Trepospira illinoisensis, ? Baylea inclinata, Baylea giffordi1, "Glabrocingulum (Ananias) talpaensis n. sp." = Ananias talpaensis, "Glabrocingulum (Glabrocingulum) grayvillense" = Glabrocingulum grayvillense, "Glabrocingulum (Glabrocingulum) globosum n. sp." = Glabrocingulum globosum, "Glyptotomaria (Dictyotomaria) sp." = Dictyotomaria, "Glyptotomaria (Dictyotomaria) ? scitula" = Dictyotomaria scitula, "Glyptotomaria (Dictyotomaria) quadrilineata" = Dictyotomaria quadrilineata, "Gosseletina spironema" = Globodoma spironema, Ptychomphalina sp.
Apachella sp. Winters 1956 snail
? Lunulazona sp. Sadlick and Nielsen 1963 snail
Luciellina ocultabanda n. sp. Kues and Batten 2001 snail
Trepospira illinoisensis Worthen 1884 snail
? Baylea inclinata Weller 1929 snail
Baylea giffordi1 Worthen 1884 snail
"Glabrocingulum (Ananias) talpaensis n. sp." = Ananias talpaensis Kues and Batten 2001 snail
"Glabrocingulum (Glabrocingulum) grayvillense" = Glabrocingulum grayvillense Norwood and Pratten 1855 snail
"Glabrocingulum (Glabrocingulum) globosum n. sp." = Glabrocingulum globosum Kues and Batten 2001 snail
"Glyptotomaria (Dictyotomaria) sp." = Dictyotomaria Knight 1945 snail
"Glyptotomaria (Dictyotomaria) ? scitula" = Dictyotomaria scitula Meek and Worthen 1860 snail
"Glyptotomaria (Dictyotomaria) quadrilineata" = Dictyotomaria quadrilineata Girty 1934 snail
"Gosseletina spironema" = Globodoma spironema Meek and Worthen 1866 snail
Ptychomphalina sp. Fischer 1885 snail
 Murchisoniina - Eotomariidae
Spiroscala cf. rockymontana, Spiroscala georgiannae n. sp., Euconospira "sp. 1", Euconospira hermosana, Euconospira turbiniformis
Spiroscala georgiannae n. sp. Kues and Batten 2001 snail
Euconospira "sp. 1" Ulrich and Scofield 1897 snail
Euconospira hermosana Girty 1934 snail
Euconospira turbiniformis Meek and Worthen 1860 snail
 Murchisoniina - Portlockiellidae
Shansiella beckwithana, Shansiella carbonaria
Shansiella beckwithana McChesney 1859 snail
Shansiella carbonaria Norwood and Pratten 1855 snail
 Murchisoniina - Porcelliidae
"Ambozone cf. gilliana" = Tomoceras gilliana
"Ambozone cf. gilliana" = Tomoceras gilliana White and St. John 1867 snail
 Murchisoniina - Phymatopleuridae
Worthenia legrandi n. sp., Worthenia speciosa, Worthenia tabulata, Phymatopleura nodosa, Paragoniozona nodolirata, Paragoniozona multilirata
Worthenia legrandi n. sp. Kues and Batten 2001 snail
Worthenia speciosa Meek and Worthen 1860 snail
Worthenia tabulata Conrad 1835 snail
Phymatopleura nodosa Girty 1912 snail
Paragoniozona nodolirata Nelson 1947 snail
Paragoniozona multilirata Nelson 1947 snail
 Euomphalina - Platyceratidae
Strophostylus girtyi
Strophostylus girtyi Knight 1934 snail
 Euomphalina - Anomphalidae
Anomphalus spp., ? Anomphalus blancus n. sp., Anomphalus verruculiferus
Anomphalus spp. Meek and Worthen 1866 snail
juveniles
? Anomphalus blancus n. sp. Kues and Batten 2001 snail
Anomphalus verruculiferus White 1881 snail
 Euomphalina - Microdomatidae
"Amaurotoma humerosa" = Knightinella humerosa, "Microdoma (Euconodoma) gavinae" = Euconodoma gavinae, Cyclites diminutus n. sp.
"Amaurotoma humerosa" = Knightinella humerosa Meek and Hayden 1858 snail
"Microdoma (Euconodoma) gavinae" = Euconodoma gavinae Kues 1990 snail
Cyclites diminutus n. sp. Kues and Batten 2001 snail
 Euomphalina - Euomphalidae
Amphiscapha ? reedsi, Amphiscapha catilloides, Euomphalus plummeri, Euomphalus serratus, Araeonema sp.
Amphiscapha ? reedsi Knight 1934 snail
Amphiscapha catilloides Conrad 1842 snail
Euomphalus plummeri Knight 1934 snail
Euomphalus serratus Knight 1934 snail
Araeonema sp. Knight 1933 snail
 Trochoidea - Eucochlidae
? Eucochlis sp., Eucochlis perminuta
? Eucochlis sp. Knight 1933 snail
Eucochlis perminuta Knight 1933 snail
 Neritoidea - Naticopsidae
Naticopsis (Jedria) ventrica, Naticopsis (Naticopsis) scintilla
Naticopsis (Jedria) ventrica Norwood and Pratten 1855 snail
 Caenogastropoda - Meekospiridae
Ceraunocochlis fulminula, Meekospira peracuta, Meekospira delgada n. sp., Meekospira choctawensis, Girtyspira minuta
Ceraunocochlis fulminula Knight 1931 snail
Meekospira peracuta Meek and Worthen 1860 snail
Meekospira delgada n. sp. Kues and Batten 2001 snail
Meekospira choctawensis Girty 1912 snail
Girtyspira minuta Stevens 1858 snail
 Caenogastropoda - Goniasmatidae
Platyzona hespera n. sp., ? Cerithioides "sp. 1", ? Cerithioides "sp. 2", "Stegocoelia (Goniasma) lasallensis" = Goniasma lasallense
Platyzona hespera n. sp. Kues and Batten 2001 snail
? Cerithioides "sp. 1" Haughton 1859 snail
? Cerithioides "sp. 2" Haughton 1859 snail
"Stegocoelia (Goniasma) lasallensis" = Goniasma lasallense Worthen 1890 snail
 Caenogastropoda - Orthonematidae
Arribazona hesperita, "Stegocoelia (Taosia) wortheni" = Taosia wortheni, "Stegocoelia (Taosia) copei" = Taosia copei, ? Orthonema "sp. 1", ? Orthonema "sp. 3", ? Orthonema "sp. 4", "Orthonema salteri" = Orthonema salteri, "Orthonema conicum" = Orthonema salteri, Orthonema semiornatum, Orthonema telescopiforme, Orthonema inornatum, Orthonema aff. ascensus, Orthonema amissidecoris, Orthonema subtaeniatum
Arribazona hesperita Kues 1990 snail
"Stegocoelia (Taosia) wortheni" = Taosia wortheni Knight 1942 snail
"Stegocoelia (Taosia) copei" = Taosia copei White 1881 snail
? Orthonema "sp. 1" Meek and Worthen 1862 snail
? Orthonema "sp. 3" Meek and Worthen 1862 snail
? Orthonema "sp. 4" Meek and Worthen 1862 snail
"Orthonema salteri" = Orthonema salteri Meek and Worthen 1860 snail
"Orthonema conicum" = Orthonema salteri Meek and Worthen 1860 snail
Orthonema semiornatum Anderson et al. 1985 snail
Orthonema telescopiforme Erwin 1988 snail
Orthonema inornatum Knight 1934 snail
Orthonema aff. ascensus Anderson et al. 1985 snail
Orthonema amissidecoris Anderson et al. 1985 snail
Orthonema subtaeniatum Geinitz 1866 snail
 Cerithioidea - Acanthonematidae
Hermosanema varium n. gen. n. sp., Bicuerda procolumnare n. gen. n. sp., ? Callispira "sp. A", Callispira quinquecostata
Hermosanema varium n. gen. n. sp. Kues and Batten 2001 snail
Bicuerda procolumnare n. gen. n. sp. Kues and Batten 2001 snail
? Callispira "sp. A" Nelson 1947 snail
Callispira quinquecostata Nelson 1947 snail
 Zygopleuroidea - Pseudozygopleuridae
Microptychis aff. cerithiformis, Microptychis turbineus, "Microptychis williamsi" = Levizygopleura williamsi, "Microptychis trochus" = Levizygopleura trochus, Cyclozyga mirabilis, "Hemizyga (Hemizyga) illineata" = Hemizyga illineata, Hemizyga (Hemizyga) "sp. 1", Hemizyga (Hemizyga) "sp. 2", Hemizyga (Hemizyga) "sp. 3", "Hemizyga (Hemizyga) larga n. sp." = Hemizyga larga, "Hemizyga (Hemizyga) attenuata" = Hemizyga attenuata, "Hemizyga (Hemizyga) ? elegans" = Hemizyga elegans, "Pseudozygopleura (Pseudozygopleura) restis" = Pseudozygopleura restis, "Pseudozygopleura (Pseudozygopleura) angusta" = Pseudozygopleura angustata, "Pseudozygopleura (Pseudozygopleura) panda" = Pseudozygopleura pandus, "Pseudozygopleura (Pseudozygopleura) scitula" = Pseudozygopleura scitula, "Pseudozygopleura (Pseudozygopleura) girtyi" = Pseudozygopleura girtyi, "Pseudozygopleura (Pseudozygopleura) semicostata" = Pseudozygopleura semicostata, "Pseudozygopleura (Pseudozygopleura) gradata" = Pseudozygopleura gradatus, "Pseudozygopleura (Pseudozygopleura) plummeri" = Pseudozygopleura plummeri, "Pseudozygopleura (Pseudozygopleura) aff. multicostata" = Pseudozygopleura multicostata, "Pseudozygopleura (Pseudozygopleura) ? pupa" = Pseudozygopleura pupa, "Pseudozygopleura (Pseudozygopleura) variegata" = Pseudozygopleura variegata, "Pseudozygopleura (Pseudozygopleura) kellettae" = Pseudozygopleura kellettae, ? Pseudozygopleura (Pseudozygopleura) "sp. 1", Pseudozygopleura (Pseudozygopleura) "sp. 2", Pseudozygopleura (Pseudozygopleura) "sp. 3", Pseudozygopleura (Pseudozygopleura) "sp. 4", Pseudozygopleura (Pseudozygopleura) "sp. 5", Pseudozygopleura (Pseudozygopleura) "sp. 6", "Pseudozygopleura (Pseudozygopleura) aff. macra" = Pseudozygopleura macra, "Hemizyga (Plocezyga) "sp. 1"" = Plocezyga, Hemizyga (Plocezyga) aff. delicata, "Hemizyga (Plocezyga) corona" = Plocezyga corona, "Hemizyga (Plocezyga) percostata" = Plocezyga percostata, "Hemizyga (Plocezyga) tenuilirata" = Plocezyga tenuilirata, "Pseudozygopleura (Stephanozyga) subnodosa" = Stephanozyga subnodosa, Pseudozygopleura (Stephanozyga) lisaspira n. sp., "Pseudozygopleura (Stephanozyga) granda n. sp." = Stephanozyga granda, Hemizyga (Hyphantozyga) knighti, Hemizyga (Hyphantozyga) fenestrata
Microptychis aff. cerithiformis Meek and Worthen 1860 snail
Microptychis turbineus Hoare and Sturgeon 1981 snail
"Microptychis williamsi" = Levizygopleura williamsi Knight 1930 snail
"Microptychis trochus" = Levizygopleura trochus Knight 1930 snail
Cyclozyga mirabilis Knight 1930 snail
"Hemizyga (Hemizyga) illineata" = Hemizyga illineata Knight 1930 snail
Hemizyga (Hemizyga) "sp. 1" Girty 1915 snail
Hemizyga (Hemizyga) "sp. 2" Girty 1915 snail
Hemizyga (Hemizyga) "sp. 3" Girty 1915 snail
"Hemizyga (Hemizyga) larga n. sp." = Hemizyga larga Kues and Batten 2001 snail
"Hemizyga (Hemizyga) attenuata" = Hemizyga attenuata Hoare and Sturgeon 1980 snail
"Hemizyga (Hemizyga) ? elegans" = Hemizyga elegans Girty 1915 snail
"Pseudozygopleura (Pseudozygopleura) restis" = Pseudozygopleura restis Knight 1930 snail
"Pseudozygopleura (Pseudozygopleura) angusta" = Pseudozygopleura angustata Hoare and Sturgeon 1985 snail
"Pseudozygopleura (Pseudozygopleura) panda" = Pseudozygopleura pandus Hoare and Sturgeon 1981 snail
"Pseudozygopleura (Pseudozygopleura) scitula" = Pseudozygopleura scitula Meek and Worthen 1860 snail
"Pseudozygopleura (Pseudozygopleura) girtyi" = Pseudozygopleura girtyi Knight 1930 snail
"Pseudozygopleura (Pseudozygopleura) semicostata" = Pseudozygopleura semicostata Meek 1871 snail
"Pseudozygopleura (Pseudozygopleura) gradata" = Pseudozygopleura gradatus Hoare and Sturgeon 1981 snail
"Pseudozygopleura (Pseudozygopleura) plummeri" = Pseudozygopleura plummeri Knight 1930 snail
"Pseudozygopleura (Pseudozygopleura) aff. multicostata" = Pseudozygopleura multicostata Meek and Worthen 1862 snail
"Pseudozygopleura (Pseudozygopleura) ? pupa" = Pseudozygopleura pupa Knight 1930 snail
"Pseudozygopleura (Pseudozygopleura) variegata" = Pseudozygopleura variegata Hoare and Sturgeon 1985 snail
"Pseudozygopleura (Pseudozygopleura) kellettae" = Pseudozygopleura kellettae Knight 1930 snail
"Pseudozygopleura (Pseudozygopleura) aff. macra" = Pseudozygopleura macra Knight 1930 snail
"Hemizyga (Plocezyga) "sp. 1"" = Plocezyga Knight 1930 snail
Hemizyga (Plocezyga) aff. delicata Hoare and Sturgeon 1980 snail
"Hemizyga (Plocezyga) corona" = Plocezyga corona Knight 1930 snail
"Hemizyga (Plocezyga) percostata" = Plocezyga percostata Knight 1930 snail
"Hemizyga (Plocezyga) tenuilirata" = Plocezyga tenuilirata Knight 1930 snail
"Pseudozygopleura (Stephanozyga) subnodosa" = Stephanozyga subnodosa Knight 1930 snail
Pseudozygopleura (Stephanozyga) lisaspira n. sp. Kues and Batten 2001 snail
"Pseudozygopleura (Stephanozyga) granda n. sp." = Stephanozyga granda Kues and Batten 2001 snail
Hemizyga (Hyphantozyga) knighti Hoare and Sturgeon 1980 snail
Hemizyga (Hyphantozyga) fenestrata Hoare and Sturgeon 1980 snail
 Cerithimorpha - Orthonemidae
"Stegocoelia (Hypergonia) hoffmani n. sp." = Hypergonia hoffmani, "Stegocoelia (Hypergonia) aff. percostata" = Hypergonia percostata, "Stegocoelia (Hypergonia) agraciada n. sp." = Hypergonia agraciada, "? Stegocoelia (? Donaldospira) sp." = Donaldospira, "Stegocoelia (Donaldospira) taosensis n. sp." = Donaldospira taosensis, Cibecuia magnum2
"Stegocoelia (Hypergonia) hoffmani n. sp." = Hypergonia hoffmani Kues and Batten 2001 snail
"Stegocoelia (Hypergonia) aff. percostata" = Hypergonia percostata Girty 1939 snail
"Stegocoelia (Hypergonia) agraciada n. sp." = Hypergonia agraciada Kues and Batten 2001 snail
"? Stegocoelia (? Donaldospira) sp." = Donaldospira Batten 1966 snail
"Stegocoelia (Donaldospira) taosensis n. sp." = Donaldospira taosensis Kues and Batten 2001 snail
Cibecuia magnum2 Mazaev 2002 snail
1Karapunar et al. 2022; 2Mazaev 2002; 3Wagner 2023