Iharkút mine, Bakony mountains (Cretaceous of Hungary)

Also known as Németbánya

Where: Veszprém, Hungary (47.2° N, 17.6° E: paleocoordinates 31.0° N, 18.6° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Csehbánya Formation, Santonian (86.3 - 83.6 Ma)

• two bone-bearing beds within section, approx. 0.5 m apart

• group of beds-level stratigraphic resolution

Environment/lithology: fine channel fill; sandy claystone and gray, argillaceous sandstone

• "deposited in an alluvial plain"; fossils "are buried in mud- and sand-filled channel deposits"

•"The palaeoenvironment is interpreted as an alluvial plain and the deposition of the breccia could be the result of the episodic filling of a channel or pond by debris flows, silt and sand" (Osi & Mindszenty, 2009).

• "The Csehbánya Formation consists of variegated clay and sandstone beds...The fauna is concentrated near the top of the outcrop in 20 cm thick sandy clay, sandstone bed...Most of the fossils derived from 5 cm thick sandy clay layer below the sandstone. Other teeth and bones occur disordered on the surface of the sandstone bed and are covered with clay less than 1 cm thick."

•"The vertebrate-bearing beds form 2–3 m thick cyclical units beginning with green clay covered by breccia containing clay clasts, sandstone and siltstone layers (Tuba et al. 2006). Most of the material, including the turtles comes from the matrix supported breccia consisting of ripped-up variegated clay clasts, dolomite pebbles, carbonized plant remains and sand forming channel-like structures with the isolated bones accumulated in lenses" (Rabi et al. 2012)

Size classes: macrofossils, mesofossils

Collected by A. Osi in 2000–2011

Collection methods: quarrying, surface (float), surface (in situ),

Primary reference: A. Osi, C.-M. Jianu, and D. B. Weishampel. 2003. Dinosaurs from the Upper Cretaceous of Hungary. In A. Petculescu & E. Stiuca (eds.), Advances in Vertebrate Paleontology: Hen to Panta. Romanian Academy, "Emil Racovita" Institute of Speleology, Bucarest 117-120 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 48722: authorized by Matthew Carrano, entered by Matthew Carrano on 30.03.2005, edited by Richard Butler, Matthew Oreska, Philip Mannion, Roger Benson, Jonathan Tennant, Terri Cleary, Patricia Holroyd and Kaitlin Maguire

Creative Commons license: CC BY (attribution)

Taxonomic list

• Fragmentary turtle plates are among the most abundant remains at the site.
 Neopterygii -
 Lepisosteiformes - Lepisosteidae
Lepisosteidae indet.10 Cuvier 1825 gar
 Temnospondyli - Albanerpetontidae
Albanerpetontidae indet.10 Fox and Naylor 1982 tetrapod
MTM V2008.22.1., V.2008.23.1., V2008.24.1., V2009.1.1., and V2010.288.1. (premaxillae); MTM V2008.28.1. and V2009.29.1. (maxillae); MTM V2008.25.1., V2008.26.1., V2008.29.1., V2009.8.1., V2009.9.1., V2009.26.1., V2009.32.1., and V2010.287.1. (dentaries) and MTM V2009.10.1. (indeterminate jaw fragment) - see Szentesi et al. (2013)
 Salientia -
Hungarobatrachus szukacsi n. gen. n. sp.23
Hungarobatrachus szukacsi n. gen. n. sp.23 Szentesi and Venczel 2010 frog
 Salientia - Discoglossidae
Bakonybatrachus fedori n. gen. n. sp.24
Bakonybatrachus fedori n. gen. n. sp.24 Szentesi and Venczel 2012 painted frog
MTM V 2010.283.1. - holotype (incomplete right ilium); referred material: fragmentary maxilla (MTM V 2009.34.1.), fragmentary angulosplenial (MTM V 2008.31.1.) and fragmentary scapula (MTM V 2008.30.1.)
 Testudines - Dortokidae
 Testudines - Bothremydidae
Bothremydidae indet.10 Baur 1891 sideneck turtle
Foxemys trabanti n. sp.21 Rabi et al. 2012 sideneck turtle
 Testudinata -
Kallokibotion sp.2 Nopcsa 1923 turtle
 Saurischia -
Titanosauriformes indet.16 Salgado et al. 1997 sauropod
MTM PAL 2017.1.1
Theropoda "indet. 3"8 Marsh 1881 theropod
"Morphotype 3"
Theropoda indet.11 Marsh 1881 theropod
MTM V 2009.48.1, sacrum fragment; MTM V 2009.47.1-6, caudal vertebra; MTM V 2009.49.1, distal left femur
Tetanurae indet.11 Gauthier 1986 tetanuran
"Morphotype 1"
 Avetheropoda -
Paraves indet.11 Sereno 1997 coelurosaur
"Morphotype 2"
 Aves -
Aves indet.1 bird
MTM V 2009.41.1, distal L femur; MTM V.2002.06, distal subadult femur; MTM V 2003.19, metatarsal III
 Enantiornithes -
Enantiornithes indet.9 Walker 1981 bird
MTM V.2002.06, fig. 2G-I, distal end of L femur; MTM V.2003.19, metatarsal III
 Enantiornithes - Avisauridae
? Bauxitornis sp.12 Dyke and Osi 2010 bird
MTM PAL 2011.20 & 21 (2 femora)
Bauxitornis mindszentyae n. gen. n. sp.1 Dyke and Osi 2010 bird
MTM V 2009.38.1, R tarsometatarsus
 Avetheropoda -
Pneumatoraptor fodori n. gen. n. sp.11
Pneumatoraptor fodori n. gen. n. sp.11 Osi et al. 2010 coelurosaur
MTM V.2008.38.1. - holotype (left scapulacoracoid)
 Saurischia - Abelisauridae
Abelisauridae indet.11 Bonaparte and Novas 1985 ceratosaur
MTM V 2008.43.1 (pedal ungual); MTM PAL 2011.17 (R femur)
 Ornithischia - Nodosauridae
Hungarosaurus sp.17 Osi 2005 ornithischian
PAL 2013.23.1, partial skull
Hungarosaurus tormai n. gen. n. sp.9 Osi 2005 ornithischian
MNI = 5
 Neornithischia -
Ajkaceratops kozmai n. gen. n. sp.14
Ajkaceratops kozmai n. gen. n. sp.14 Osi et al. 2010 ornithischian
MTM V 2009.192.1 (holotype), fused premaxillae and rostral bones. MTM V2009.193.1, V2009.194.1, V2009.195.1, V2009.196.1, predentaries
 Neornithischia - Rhabdodontidae
Rhabdodontidae indet. Weishampel et al. 2003 rhabdodontid
Mochlodon vorosi n. sp.18 Osi et al. 2012 rhabdodontid
Left complete dentary with four broken teeth (MTM V 2010.105.1; holotype); left postorbital (MTM 2012.14.1); two right quadrates (MTM V 2010.110.1, V 2010.111.1), two left (MTM V 2010.105.1., 2012.15.1) and two right (MTM V 2010.107.1., V 2010.109.1.) dentaries, all four of which are almost complete, six fragmentary dentaries (MTM V 2010 106.1, V 2010 107.1, V 2010 108.1, V 2010 109.1, V 2010.112.1, 2012.16.1), 15 maxillary and 23 dentary teeth (MTM V 2000.01., V 2000.32., V 2000.33., V 2003.10., V 01.161., V 2003.14,– V.2003.16, V 01.64., 2012.17.1, 2012.18.1), isolated cervical (MTM 2012.19.1), dorsal (MTM 2010.118.1.), and caudal (MTM 2012.20.1, 2012.21.1) vertebrae, almost complete but compressed sacrum (MTM V 2010.121.1.), three coracoids (MTM V 01.53., V 2010.122.1., V 2010.123.1.), one fragmentary scapula (MTM 2012.22.1), one fragmentary (MTM 2012.23.1) and one complete humerus (MTM V 2010.128.1.), one complete ulna (MTM 2012.24.1), two almost complete femora (MTM V 01.225., V 2010.126.1.), one fragmentary femur (MTM 2012.25.1), one complete tibia (MTM V 2010.127.1.), two fragmentary tibiae (MTM V 01.101., 2012.26.1), and two phalanges (MTM 2012.27.1, 2012.28.1).
 Pterosauria -
Pterodactyloidea indet.13 pterosaur
MTM V 2010.98.1 (L articular region of lower jaw), MTM V 2010.99.1 (MC IV)
 Pterosauria - Azhdarchidae
Bakonydraco galaczi n. gen. n. sp.19 Osi et al. 2005 pterosaur
MTM Gyn/3 (holotype), nearly complete mandible. MTM Gyn/4, 21 dentary symphyseal fragments; MTM V 2010.80.1 (frag. premaxilla), 2010.74.1–74.22 (dentary symphyseal frags.)
 Loricata -
Doratodon carcharidens20 Bunzel 1871 crocodilian
Single fragmentary dentary and several isolated teeth
 Crocodylia -
Alligatoroidea indet.10 crocodilian
A few isolated skull fragments and hundreds of teeth
 Loricata - Hylaeochampsidae
Iharkutosuchus makadii15 Osi et al. 2007 crocodilian
 Squamata - Teiidae
Teiidae indet.10 Gray 1827 squamates
 Squamata -
Pannoniasaurus inexpectatus n. gen. n. sp.6 Makádi et al. 2012 squamates
 Squamata - Polyglyphanodontidae
Bicuspidon aff. hatzegiensis3 Folie and Codrea 2005 squamates
MTM V.01.24., left dentary fragment; MTM V.01.27., right lower jaw fragment; MTM 2006.108.1, left dentary; MTM 2006.109.1, left dentary fragment; MTM 2006.110.1, dentary fragment; MTM 2006.111.1, left dentary; MTM 2006.112.1, left dentary; MTM 2006.114.1, jaw fragment
Distortodon rhomboideus n. gen. n. sp.4 Makadi 2013 squamates
Holotype. MTM PAL2012.31.1. partial right maxilla; MTM PAL2012.32.1. right dentary (Fig. 3) and MTM PAL2012.33.1. partial left dentary
 Squamata -
Lacertilia indet. lizard
 Squamata - Chamopsiidae
Pelsochamops infrequens n. gen. n. sp.5
Pelsochamops infrequens n. gen. n. sp.5 Makadi 2013 lizard
MTM 2006.106.1 (holotype), partial left mandible; MTM PAL 2013.24.1, right dentary fragment; MTM PAL 2013.25.1, left dentary fragment
 Squamata -
Chromatogenys tiliquoides n. gen. n. sp.7
Chromatogenys tiliquoides n. gen. n. sp.7 Makadi and Nydam 2015 lizard
MTM V 2010.129.1 (holotype), partial mandible
Plantae indet.9 Haeckel 1866
"carbonized plant fragments"