La Cantalera 1, Josa (Cretaceous of Spain)

Where: Aragón, Spain (41.0° N, 0.8° W: paleocoordinates 30.9° N, 9.4° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Atopochara trivolvis triquetra other zone, Cabezo Gordo Member (Blesa Formation), Early/Lower Barremian (130.0 - 125.5 Ma)

• "Calizas y Margas de Blesa"; facies C

•Lowest part of the lower Blesa Formation (Badiola et al 2012)

• bed-level stratigraphic resolution

Environment/lithology: mire or swamp; paleosol/pedogenic, gray, carbonaceous claystone

• "non-permanent water marshy area"
• "a wide outcrop of grey clays with very abundant vegetal remains, carbonate nodules formed in paleosols, and reelaborated marine fossils. The site was formed in a palustrine environment with no permanent water and abundant vegetation in the surrounding area"

Size classes: macrofossils, mesofossils, microfossils

Collected in 1997–2000

Collection methods: quarrying, surface (float), surface (in situ), mechanical, sieve,

Primary reference: J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1997. Primera evidencia de un área de alimentación de dinosaurios herbívoros en el Cretácico Inferior de España (Teruel) [First evidence of a feeding area of herbivorous dinosaurs in the Lower Cretaceous of Spain (Teruel)]. Monografias de la Academia de Ciencias Exactas, Fisicas, Quimicas y Naturales de Zaragoza 10:1-48 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 53921: authorized by Matthew Carrano, entered by Kaitlin Maguire on 16.09.2005, edited by Matthew Carrano, Roger Benson and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• Among the microfossils, ostracods and charophytes are abundant, forming the greater part of the residue of the screen washing over 50 micro-m. Also frequent are plant fragments, microvertebraes remains (mainly teeth), and continental gastropods.
unclassified
  -
 Charales - Clavatoraceae
Pseudoglobator sp. Grambast 1969
Atopochara trivolvis Peck 1938
A. trivolvis triquetra
Ostracoda
 Podocopida - Cyprididae
Cypridea soriana ostracod
C. (Ulwellia) soriana
Gastropoda
 Pulmonata -
 Architaenioglossa - Viviparidae
Viviparidae indet. Gray 1847 snail
Reptilia
 Dinosauria -
Dinosauria indet. dinosaur
including eggshell, coprolites
 Saurischia -
Somphospondyli indet.8 Wilson and Sereno 1998 sauropod
MPZ 97/464, 465
 Ornithischia -
Thyreophora indet.14 ornithischian
Ankylosauria indet.13 Osborn 1923 ankylosaur
MPZ 2018/503, MPZ 2018/504, MPZ 2018/505, MPZ 2018/506, MPZ 2018/507, MPZ 2018/508, MPZ 2018/509, MPZ 2018/510-512
 Ornithischia - Polacanthidae
Polacanthidae indet.7 Wieland 1911 ankylosaur
MPZ 2002/926, 297
 Ornithischia -
Ornithopoda indet. Marsh 1881 ornithopod
MPZ 97/477-484, 504-518; including (possibly) coprolites
Ornithopoda indet.13 Marsh 1881 ornithopod
MPZ 2018/491, MPZ 2018/492, MPZ 2018/493, MPZ 2018/494, MPZ 2018/495, MPZ 2018/497, MPZ 2018/498, MPZ 2018/499, MPZ 2018/500, MPZ 2018/501, MPZ 2018/502
 Ornithischia - Rhabdodontidae
? Rhabdodontidae indet.5 Weishampel et al. 2003 rhabdodontid
Tooth, CAN1-1616
 Ornithischia -
? Hadrosauroidea indet.5 Huene 1954 ornithopod
MPZ 97/477, CAN1-925
 Ornithischia - Spheroolithidae
cf. Spheroolithidae "indet. 1"5 Zhao 1979 ornithopod
CAN1-1622
cf. Spheroolithidae "indet. 2"5 Zhao 1979 ornithopod
CAN1-1621
"Macroolithus turolensis" = Guegoolithus turolensis5 Amo Sanjuán et al. 2000 ornithopod
CAN1-1617, MPZ 2013/ 868 to MPZ 2013/1200
 Ornithischia -
Iguanodontoidea indet.9 Hay 1902 ornithopod
MPZ 97/473-476, 485-490, 502-503
Iguanodontoidea indet.9 Hay 1902 ornithopod
MPZ 97/469-472
 Ornithischia - Hypsilophodontidae
Hypsilophodontidae indet. hypsilophodontid
MPZ 97/491-501
  - Prismatoolithidae
Prismatoolithus sp.5 Zhao and Li 1993 dinosaur
CAN1-1619
Trigonoolithus amoae n. gen. n. sp.11 Moreno-Azanza et al. 2014 dinosaur
CAN1-1620; MPZ 2012/428 to MPZ 2012/868
 Theropoda -
Theropoda "indet. A" theropod
MPZ 97/466-467
"Prodeinodon" sp.15 Osborn 1924 tetanuran theropod
Teeth
Coelurosauria indet.14 coelurosaur
Maniraptora "indet. 1"5 Gauthier 1986 maniraptoran
Teeth, CAN1-1031
Maniraptora "indet. 2"5 Gauthier 1986 maniraptoran
Teeth, CAN1-1043
Maniraptora "indet. 3"5 Gauthier 1986 maniraptoran
Teeth, MPZ2003/232
? Avialae indet.5 Gauthier 1986 avialan
aff. Paronychodon sp.5 Cope 1876 maniraptoran
Teeth, CAN1-1035
 Theropoda - Dromaeosauridae
Dromaeosauridae indet.14 Colbert and Russell 1969 maniraptoran
Velociraptorinae indet.5 Barsbold 1983 maniraptoran
Teeth, CAN1-1029
Dromaeosaurinae "indet. 1"5 Matthew and Brown 1922 maniraptoran
Teeth, CAN1-979
Dromaeosaurinae "indet. 3"5 Matthew and Brown 1922 maniraptoran
Teeth, CAN1-1058
 Theropoda - Spinosauridae
? Spinosaurinae indet.10, "Baryonychidae indet." = Baryonychinae6, "Baryonychinae indet." = Baryonychinae10, "Baryonychinae indet." = Baryonychinae10
? Spinosaurinae indet.10 Sereno et al. 1998 tetanuran theropod
CAN1 960, CAN1 968, CAN1 973, CAN1 990; teeth
"Baryonychidae indet." = Baryonychinae6 Charig and Milner 1986 tetanuran theropod
CAN1 953, 957–959, 977, 983-984, 999, 1616; CAN1 974; MPZ 2001/207- 208
"Baryonychinae indet." = Baryonychinae10 Charig and Milner 1986 tetanuran theropod
MPZ 97/468
"Baryonychinae indet." = Baryonychinae10 Charig and Milner 1986 tetanuran theropod
Teeth, CAN1-1001
 Theropoda - Elongatoolithidae
Elongatoolithidae indet.14 Zhao 1975 theropod
CAN1-1618
 Theropoda - Carcharodontosauridae
? Carcharodontosauridae indet.5 Stromer 1931 allosauroid
Teeth, CAN1-963
 Pterosauria - Istiodactylidae
? Istiodactylidae indet.5 Howse et al. 2001 pterosaur
 Pterosauria - Ornithocheiridae
? Ornithocheiridae indet.5 pterosaur
Teeth
 Loricata - Krokolithidae
Krokolithes sp.5 Hirsch 1985 crocodile eggshell
CAN1-1623
 Loricata -
cf. Lisboasaurus sp.5 Seiffert 1975 crocodilian
Teeth
 Loricata - Atoposauridae
aff. Theriosuchus sp.16 Owen 1878 crocodilian
 Loricata - Goniopholididae
"Goniopholidae indet." = Goniopholididae
"Goniopholidae indet." = Goniopholididae Cope 1875 crocodilian
Teeth
 Loricata - Bernissartiidae
Bernissartiidae indet. Dollo 1883 crocodilian
Teeth
 Loricata -
Mycomorphoolithus kohringi n. gen. n. sp.12
Mycomorphoolithus kohringi n. gen. n. sp.12 Moreno-Azanza et al. 2015 crocodilian
MPZ2014/2–MPZ2014/150, eggshell fragments
 Testudinata - Pleurosternidae
? Pleurosternidae indet.7 Cope 1868 turtle
? Pleurosternidae indet.5 Cope 1868 turtle
"Chelonii indet."
 Squamata - Paramacellodidae
? Paramacellodidae indet.5 Estes 1983 squamates
Mammalia
 Multituberculata -
Plagiaulacida indet.1 Ameghino 1889 multituberculate
Fragment of an I2, FCPT (CAN 1 ⁄ 939), and fragment of a ?ix, FCPT (CAN 1 ⁄ 938) (Badiola et al 2008)
Plagiaulacida indet.2 Ameghino 1889 multituberculate
 Multituberculata - Albionbaataridae
Albionbaataridae indet.7 Kielan-Jaworowska and Ensom 1994 multituberculate
 Multituberculata - Pinheirodontidae
Cantalera abadi n. gen. n. sp.2
Cantalera abadi n. gen. n. sp.2 Badiola et al. 2008 multituberculate
 Multituberculata -
Iberica hahni n. gen. n. sp.3, ? Iberica hahni3
Iberica hahni n. gen. n. sp.3 Badiola et al. 2011 multituberculate
FCPT (CAN 1/936), left P1/3 (holotype)
? Iberica hahni3 Badiola et al. 2011 multituberculate
FCPT (CAN 1/937), fragment of a right p4
 Multituberculata - Eobaataridae
Eobaatar sp.2 Kielan-Jaworowska et al. 1987 multituberculate
Amphibia
 Temnospondyli - Albanerpetontidae
Albanerpetontidae indet.5 Fox and Naylor 1982 tetrapod
Two right humeri (incl. CAN 1-1626) and four dentaries (incl. CAN 1-1627)
 Salientia - Discoglossidae
Discoglossidae indet.5 Günther 1859 painted frog
Actinopteri
 Teleostei -
Teleostei indet.5 Müller 1846