James T. Thomas estate (Miocene of the United States)

Where: Charles County, Maryland (38.5° N, 76.7° W: paleocoordinates 38.7° N, 73.9° W)

• coordinate estimated from map

When: Calvert Beach Member (Calvert Formation), Serravallian (13.8 - 11.6 Ma)

• Cope refers to the beds as "Yorktown" but they are much more likely to be Calvert Formation since the Yorktown is not thought to crop out in Maryland, DN5. Zones 14-16A are regarded as Serravallian in age.

•Age and Member after Weems and Edwards (2007).

• bed-level stratigraphic resolution

Environment/lithology: offshore shelf; lithology not reported

• sparse molluscs, low in diversity; subtropical environment in embayment and dry onshore; area not far from shore, bay, short-term eustatic transgression events, long-term regression trend

Size classes: macrofossils, mesofossils

Preservation: original phosphate

Primary reference: E. D. Cope. 1867. An addition to the vertebrate fauna of the Miocene period, with a synopsis of the extinct Cetacea of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 19(4):138-157 [M. Uhen/M. Uhen/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 55396: authorized by Mark Uhen, entered by Mark Uhen on 14.10.2005, edited by Austin Hendy

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
 Life -
Aves
 Procellariiformes - Procellariidae
Puffinus sp.9 Brisson 1760 shearwater
Puffinus conradi n. sp.7 Marsh 1870 shearwater
Reptilia
 Crocodylia -
"Thecachampsa contusor n. sp." = Thecachampsa antiquus, Thecachampsa sericodon n. sp.
"Thecachampsa contusor n. sp." = Thecachampsa antiquus Leidy 1852 crocodilian
ANSP 9152 - type
Thecachampsa sericodon n. sp. Cope 1867 crocodilian
ANSP 9443-9445 - type
 Testudines - Cheloniidae
Chelone sp. Linnaeus 1758 sea turtle
 Testudines - Pantrionychidae
Trionyx cellulosus Cope 1868 turtle
Mammalia
 Cetacea - Platanistidae
Zarhachis flagellator n. gen. n. sp.3
Zarhachis flagellator n. gen. n. sp.3 Cope 1868 toothed whale
 Cetacea -
Priscodelphinus atropius n. sp.3, Priscodelphinus stenus n. sp.3, Priscodelphinus acutidens n. sp., Priscodelphinus spinosus n. sp.3, Cetophis heteroclitus n. gen. n. sp.3, "Delphinapterus ruschenbergeri n. sp." = Tretosphys ruschenbergeri3, "Delphinapterus lacertosus n. sp." = Tretosphys lacertosus3, "Delphinapterus hawkinsii n. sp." = Tretosphys lacertosus3, "Delphinapterus gabbii n. sp." = Tretosphys gabbii3, Ixacanthus coelospondylus n. gen. n. sp.3, Rhabdosteus latiradix n. gen. n. sp.2, "Squalodon mento n. sp." = Delphinodon mento2
Priscodelphinus atropius n. sp.3 Cope 1868 toothed whale
Priscodelphinus stenus n. sp.3 Cope 1868 toothed whale
Priscodelphinus acutidens n. sp. Cope 1867 toothed whale
Priscodelphinus spinosus n. sp.3 Cope 1868 toothed whale
Cetophis heteroclitus n. gen. n. sp.3 Cope 1868 toothed whale
"Delphinapterus ruschenbergeri n. sp." = Tretosphys ruschenbergeri3 Cope 1868 toothed whale
"Delphinapterus lacertosus n. sp." = Tretosphys lacertosus3 Cope 1868 toothed whale
"Delphinapterus hawkinsii n. sp." = Tretosphys lacertosus3 Cope 1868 toothed whale
"Delphinapterus gabbii n. sp." = Tretosphys gabbii3 Cope 1868 toothed whale
Ixacanthus coelospondylus n. gen. n. sp.3 Cope 1868 toothed whale
Rhabdosteus latiradix n. gen. n. sp.2 Cope 1867 toothed whale
"Squalodon mento n. sp." = Delphinodon mento2 Cope 1867 toothed whale
It is not clear which locality Squalodon mento is actually from.
 Cetacea - Physeteridae
Orycterocetus crocodilinus Cope 1867 sperm whale
 Cetacea -
Squalodon wymanii n. sp., "Balaenoptera pusilla n. sp." = Eschrichtius pusillus4
Squalodon wymanii n. sp. Cope 1867 toothed whale
it is not clear which locality Squalodon wymanii is from. Only described as Miocene of Charles County, MD.
"Balaenoptera pusilla n. sp." = Eschrichtius pusillus4 Cope 1868 whale
 Cetacea - Cetotheriidae
"Delphinapterus tyrannus n. sp." = Cetotherium parvum3
"Delphinapterus tyrannus n. sp." = Cetotherium parvum3 Trouessart 1898 whale
 Cetacea - Balaenopteridae
? Balaenoptera cephalus8 Cope 1867 rorqual
 Cetacea -
"Megaptera expansa n. sp." = Eschrichtius expansus3
"Megaptera expansa n. sp." = Eschrichtius expansus3 Cope 1868 whale
Actinopteri
 Scombriformes -
"Sphyraena speciosa" = Sphyraenodus speciosus
"Sphyraena speciosa" = Sphyraenodus speciosus Leidy 1856
Chondrichthyes
 Carcharhiniformes - Galeocerdidae
"Galeocerdo contortus" = Physogaleus contortus Gibbes 1847 ground shark
Galeocerdo sp. Müller and Henle 1837 tiger shark
Galeocerdo latidens Agassiz 1835 tiger shark
Galeocerdo egertoni Agassiz 1843 tiger shark
Galeocerdo aduncus Agassiz 1835 tiger shark
 Carcharhiniformes - Sphyrnidae
Sphyrna sp., Sphyrna magna, "Galeocerdo laevissimus n. sp." = Sphyrna laevissima
Sphyrna sp. Rafinesque 1810 hammerhead shark
Sphyrna magna Cope 1868 hammerhead shark
"Galeocerdo laevissimus n. sp." = Sphyrna laevissima Cope 1867 hammerhead shark
 Carcharhiniformes - Hemigaleidae
Hemipristis serra Agassiz 1835 ground shark
 Lamniformes - Otodontidae
Otodus obliquus, "Carcharodon megalodon" = Otodus megalodon, "Carcharodon ? angustidens" = Otodus (Carcharocles) angustidens, "Otodus appendiculatus" = Cretalamna appendiculata
Otodus obliquus Agassiz 1838 mackerel shark
"Carcharodon megalodon" = Otodus megalodon Agassiz 1835 megalodon
"Carcharodon ? angustidens" = Otodus (Carcharocles) angustidens Agassiz 1835 mackerel shark
"Otodus appendiculatus" = Cretalamna appendiculata Agassiz 1835 mackerel shark
 Lamniformes - Odontaspididae
"Lamna cuspidata" = Araloselachus cuspidatus, "Lamna denticulata" = Araloselachus cuspidatus
"Lamna cuspidata" = Araloselachus cuspidatus Agassiz 1843 sand tiger shark
"Lamna denticulata" = Araloselachus cuspidatus Agassiz 1843 sand tiger shark
 Lamniformes - Mitsukurinidae
"Lamna elegans" = Striatolamia macrota, "Lamna ? subulata" = Scapanorhynchus subulatus
"Lamna elegans" = Striatolamia macrota Agassiz 1838 goblin shark
"Lamna ? subulata" = Scapanorhynchus subulatus Agassiz 1843 goblin shark
 Lamniformes - Carchariidae
"Lamna hopei" = Carcharias hopei
"Lamna hopei" = Carcharias hopei Agassiz 1843 sand shark
 Hexanchiformes - Hexanchidae
"Notidanus plectrodon n. sp." = Notorynchus cepedianus, Notidanus primigeneus
"Notidanus plectrodon n. sp." = Notorynchus cepedianus Peron 1807 cow shark
 Heterodontiformes - Heterodontidae
"Sphyrna prisca" = Cestracion prisca
"Sphyrna prisca" = Cestracion prisca Agassiz 1843 bullhead shark
 Myliobatiformes - Myliobatidae
Myliobatis gigas Cope 1868 eagle ray
Myliobatis vicomicanus Cope 1868 eagle ray
Myliobatis pachyodon Cope 1868 eagle ray
Aetobatus arcuatus Agassiz 1843 eagle ray
[entered as Aetobatis arcuatus]
"Aetobatis arcuatus" = Aetobatus irregularis6 Agassiz 1843 eagle ray
Aetobatis profundus n. sp. Cope 1867 eagle ray
 Myliobatiformes - Dasyatidae
"Raja dux" = Trygon dux
"Raja dux" = Trygon dux Cope 1867 whiptail stingray
Bivalvia
 Pectinida - Pectinoidae
"Pecten jeffersonius" = Chesapecten jeffersonius, "Pecten madisonius" = Chesapecten madisonius, Pecten humphreysii, "Pecten virginianus" = Placopecten virginiana, "Pecten clintonius" = Placopecten clintonius, "Pecten marylandicus" = Placopecten marylandica, "Pecten cerinus n. sp." = Eburneopecten cerinus
"Pecten jeffersonius" = Chesapecten jeffersonius Say 1824 scallop
"Pecten madisonius" = Chesapecten madisonius Say 1824 scallop
Pecten humphreysii Conrad 1842 scallop
"Pecten virginianus" = Placopecten virginiana Conrad 1840 scallop
"Pecten clintonius" = Placopecten clintonius Say 1824 scallop
"Pecten marylandicus" = Placopecten marylandica Wagner 1839 scallop
"Pecten cerinus n. sp." = Eburneopecten cerinus Conrad 1869 scallop
 Pectinida - Anomiidae
 Ostreida - Pinnidae
Pinna sp. Linnaeus 1758 oyster
 Ostreida - Malleidae
 Ostreida - Ostreidae
Ostreidae indet.1 Rafinesque 1815 oyster
Ostrea trachydiscus n. sp. oyster
 Arcida - Arcidae
 Arcida - Glycymerididae
"Axinaea lentiformis" = Glycymeris, Glycymeris americana5
"Axinaea lentiformis" = Glycymeris clam
Glycymeris americana5 Defrance 1826 clam
 Arcida - Noetiidae
 Lucinida - Lucinidae
"Lucina contracta" = Lucinoma contracta, "Lucina foremani" = Stewartia foremani, "Lucina americana" = Divalinga quadrisulcata
"Lucina contracta" = Lucinoma contracta Say 1824 clam
"Lucina foremani" = Stewartia foremani Conrad 1841 clam
"Lucina americana" = Divalinga quadrisulcata d'Orbigny 1846 clam
 Cardiida - Chamidae
Chama corticosa jewel box
 Cardiida - Glossidae
"Isocardia fraterna" = Glossus fraterna, Isocardia markoii
"Isocardia fraterna" = Glossus fraterna Say 1824 clam
 Cardiida - Veneridae
"Dione staminaea" = Hysteroconcha, "Dione marylandica" = Hysteroconcha, Mercenaria staminea, Mercenaria cuneata n. sp., Dosinia acetabulum
"Dione staminaea" = Hysteroconcha venus clam
"Dione marylandica" = Hysteroconcha venus clam
Mercenaria cuneata n. sp. Conrad 1869 quahog
Dosinia acetabulum Conrad 1832 venus clam
 Cardiida - Cardiidae
"Cardium laqueatum" = Chesacardium laqueatum
"Cardium laqueatum" = Chesacardium laqueatum Conrad 1830 cockle
 Cardiida - Tellinidae
"Metis biplicata" = Florimetis biplicata
"Metis biplicata" = Florimetis biplicata Conrad 1834 tellin clam
 Pholadida - Corbulidae
"Corbula idonea" = Bicorbula idonea, Corbula pectorosa n. sp.
"Corbula idonea" = Bicorbula idonea Conrad 1833 clam
Corbula pectorosa n. sp. clam
 Carditida - Crassatellidae
"Crassatella marylandica" = Marvacrassatella marylandica, "Crassatella melina" = Marvacrassatella melina
"Crassatella marylandica" = Marvacrassatella marylandica Conrad 1832 clam
"Crassatella melina" = Marvacrassatella melina Conrad 1832 clam
 Carditida - Astartidae
Astarte cuneiformis Conrad 1840 clam
Astarte perplana Conrad 1840 clam
Astarte undata5 Gould 1841 clam
 Carditida - Carditidae
"Carditamera protracta" = Cardita protracta, Cyclocardia granulata
"Carditamera protracta" = Cardita protracta Conrad 1843 clam
Cyclocardia granulata Say 1824 clam
Gastropoda
 Lepetellida - Fissurellidae
"Fissurella redimicula" = Diodora redimicula
"Fissurella redimicula" = Diodora redimicula Say 1824 snail
 Sorbeoconcha - Ranellidae
"Bursa centrosa n. sp." = Tritonium centrosum
"Bursa centrosa n. sp." = Tritonium centrosum Cope 1867 triton
 Neogastropoda - Muricidae
Ecphora quadricostata Conrad 1830 murex snail
 Sorbeoconcha - Naticidae
 Epitonioidea - Epitoniidae
Scala pachypleura Conrad 1841 wentletrap
 Cerithioidea - Turritellidae
Turritella indenta Conrad 1841 turret shell
"Turritella alticostata" = Turritella (Turritella) alticostata Conrad 1834 turret shell
Craniata
 Craniida - Craniidae
"Orbicula lububris" = Neocrania
"Orbicula lububris" = Neocrania
Thecostraca
 Balanomorpha - Balanidae
Balanus vulcanellus n. sp., "Balanus proteus" = Chesaconcavus proteus
Balanus vulcanellus n. sp. barnacle
"Balanus proteus" = Chesaconcavus proteus Conrad 1834 barnacle
Anthozoa
 Scleractinia - Acroporidae
"Madrepora palmata" = Acropora palmata
"Madrepora palmata" = Acropora palmata Lamarck 1816 stony coral
Dinophyceae
 Peridiniales - Protoperidiniaceae
 Peridiniales - Deflandreaceae
 Peridiniales -
  -
Spiniferites sp.8 Mantell 1850
Spiniferites mirabilis8 Sarjeant 1970
 Gonyaulacales -
 Gonyaulacales - Gonyaulacaceae
 Gonyaulacales -
Reticulatosphaera actinocoronata8 Bujak and Matsuoka 1986
  -
 Dinoflagellata -
Pentadinium cf. latincinctum8
subspecies granulatum
Crassatella undulata
  -
"Crassatella undulata" = Kalolophus antillarum
"Crassatella undulata" = Kalolophus antillarum Reeve 1842