PBDB Collection

Parana, Pueblo Brugo to Diamante, Ituzaingo Fm. (Neogene of Argentina)

Also known as “Conglomerado osífero” (= “Mesopotamiense”, sensu Frenguelli, 1920)

Where: Entre Rios, Argentina (31.5° S, 60.2° W: paleocoordinates 31.6° S, 58.1° W)

• coordinate estimated from map

• local area-level geographic resolution

When: Huayquerian mammal zone, Ituzaingo Formation, Huayquerian (8.0 - 5.0 Ma)

• Schmidt 2015: fossils from the base of the Ituzaingo Fm.

Environment/lithology: marine; sandstone and mudstone

Size classes: macrofossils, mesofossils

Preservation: original phosphate

Primary reference: A. L. Cione, M. M. Azpelicueta, M. Bond, A. A. Carlini, J. R. Casciotta, M. A. Cozzuol, M. Fuente, Z. Gasparini, F. J. Goin, J. Noriega, G. J. Scillato-Yane, L. Soibelzon, E. P. Tonni, D. H. Verzi, and M. G. Vucetich. 2000. Miocene vertebrates from Entre Rios province, eastern Argentina. Serie Correlacion Geologica 14:191-237 [M. Uhen/M. Uhen/O. Alfaro]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 55600: authorized by Mark Uhen, entered by Mark Uhen on 01.11.2005, edited by Philip Mannion, Juan Carrillo, Miranta Kouvari, Evangelos Vlachos, Grace Varnham and Kateryn Pino

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

unclassified
-	Macrahinga paranensis, Dentrocheninae indet., Mycteriini indet. Macrahinga paranensis Dentrocheninae indet. Mycteriini indet.
Aves
Phoenicopteriformes - Phoenicopteridae	Phoenicopteridae indet. Phoenicopteridae indet. Bonaparte 1838 flamingo
Phoenicopteriformes - Palaelodidae	Megapaloelodus sp. Megapaloelodus sp. Miller 1944 swimming flamingo
Gruiformes - Phorusrhacidae	"Phororhacidae indet." = Phorusrhacidae, "? Onactornis sp." = Devincenzia, Andalgalornis sp. "Phororhacidae indet." = Phorusrhacidae Ameghino 1889 terror bird "? Onactornis sp." = Devincenzia Kraglievich 1932 terror bird Andalgalornis sp. Patterson and Kraglievich 1960 terror bird
Gruiformes - Rallidae	Rallidae indet. Rallidae indet. Vigors 1825 rail
Rheiformes - Rheidae	Rheidae indet. Rheidae indet. Bonaparte 1849 rhea
Reptilia
Loricata -	Crocodylia indet. Crocodylia indet. crocodilian
Crocodylia - Gavialidae	Gryposuchus neogaeus Gryposuchus neogaeus Rusconi 1933 crocodilian
Crocodylia - Alligatoridae	"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi¹¹, Caiman jacare, Caiman australis, Caiman cf. yacare¹, Caiman latirostris, "Caiman gasparinae n. sp." = Paranasuchus gasparinae³, Paranacaiman bravardi n. gen. n. sp.² "Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi¹¹ Bocquentin Villanueva 1984 crocodilian MLP 73-IV-15-8 - holotype (posterior portion of skull); hypodigm: MLP 73-IV-15-9 (posterior portion of skull) Caiman jacare caiman Caiman australis Bravard 1858 caiman Caiman cf. yacare¹ Daudin 1802 caiman MLP 73-IV-15-5, MLP 73-IV-15-6, MLP 73-IV-5-17, MACN PV 5417 Caiman latirostris Daudin 1801 broud-snouted caiman "Caiman gasparinae n. sp." = Paranasuchus gasparinae³ Bona and Carabajal 2013 crocodilian MLP-73-IV-15-1 - holotype (skull fragments represented by a rostrum with articulated fragments of premaxillae, maxillae, nasals, left lacrymal and a partial braincase lacking the basicranium); referred material - MACN PV 5555 (fragment of right premaxilla) Paranacaiman bravardi n. gen. n. sp.² Bona et al. 2024 crocodilian MACN-PV 13551
Squamata -	Tupinambis cf. merianae Tupinambis cf. merianae tegu
Testudines - Cheloniidae	Cheloniidae indet.¹⁰ Cheloniidae indet.¹⁰ Bonaparte 1832 sea turtle MACN Pv 14224, nearly complete right mandible
Testudines - Chelidae	Phrynops cf. geoffroanus Phrynops cf. geoffroanus Schweigger 1812 side-necked turtle of Geoffroy
Mammalia
Cingulata -	"Berthawyleria sp." = Pseudoplohophorus "Berthawyleria sp." = Pseudoplohophorus Castellanos 1926 edentate
Cingulata - Panochthidae	Hoplophorus verus²³ Hoplophorus verus²³ Ameghino 1889 edentate
Cingulata - Glyptodontidae	Plohophorus paranensis, Eleutherocercus sp.²⁰, Comaphorus concisus, Paraglyptodon paranensis²³ Plohophorus paranensis Ameghino 1891 glyptodon Eleutherocercus sp.²⁰ Koken 1888 glyptodon Comaphorus concisus Ameghino 1886 glyptodon Paraglyptodon paranensis²³ Castellanos 1953 glyptodon
Cingulata - Pampatheriidae	Kraglievichia paranense, "Scirrotherium carinatum n. sp." = Kraglievichia carinatum²³ Kraglievichia paranense Ameghino 1883 edentate "Scirrotherium carinatum n. sp." = Kraglievichia carinatum²³ Gois et al. 2013 edentate
Cingulata -	Protoglyptodon primiformis, Palaehoplophorus antiquus, Chlamyphractus pressulus, Parahoplophorus paranensis, ? Trachycalyptus cingulatus, Pseudoeuryurus lelongianus, Urotherium interundatum Protoglyptodon primiformis Ameghino 1885 edentate Palaehoplophorus antiquus Ameghino 1883 edentate Chlamyphractus pressulus Ameghino 1885 edentate Parahoplophorus paranensis Ameghino 1883 edentate ? Trachycalyptus cingulatus Ameghino 1889 edentate Pseudoeuryurus lelongianus Ameghino 1889 edentate Urotherium interundatum Ameghino 1885 edentate
Cingulata - Dasypodidae	Chasicotatus spinozai²³, Macroeuphractus retusus²², Proeuphractus limpidus, ? Zaedyus sp., Dasypus neogaeus Chasicotatus spinozai²³ Scillato-Yané et al. 2010 armadillo Macroeuphractus retusus²² Ameghino 1887 armadillo but also ref 15016 Proeuphractus limpidus Ameghino 1886 armadillo ? Zaedyus sp. Ameghino 1889 armadillo Dasypus neogaeus Ameghino 1891 armadillo
Megatherioidea - Megalonychidae	"Pliomorphus mutilatus" = Pliomorphus mutilatus, "Pliomorphus ameghinoi" = Pliomorphus mutilatus, "Pliomorphus brevis" = Pliomorphus mutilatus, Pliomorphus robustus, "Ortotherium laticurvatum" = Ortotherium laticurvatum, "Ortotherium scrofum" = Ortotherium laticurvatum, "Amphiocnus seneus" = Ortotherium laticurvatum, "Ortotherium schlosseri" = Ortotherium laticurvatum, Ortotherium robustum, Protomegalonyx praecursor, Protomegalonyx doellojuradoi, Amphiocnus paranense, Megalonychops primigenius, Torcellia paranense, "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum "Pliomorphus mutilatus" = Pliomorphus mutilatus Ameghino 1885 edentate "Pliomorphus ameghinoi" = Pliomorphus mutilatus Ameghino 1885 edentate "Pliomorphus brevis" = Pliomorphus mutilatus Ameghino 1885 edentate Pliomorphus robustus Ameghino 1885 edentate "Ortotherium laticurvatum" = Ortotherium laticurvatum Ameghino 1885 edentate "Ortotherium scrofum" = Ortotherium laticurvatum Ameghino 1885 edentate "Amphiocnus seneus" = Ortotherium laticurvatum Ameghino 1885 edentate "Ortotherium schlosseri" = Ortotherium laticurvatum Ameghino 1885 edentate Ortotherium robustum Ameghino 1881 edentate Protomegalonyx praecursor Kraglievich 1925 edentate Protomegalonyx doellojuradoi Kraglievich 1925 edentate Amphiocnus paranense Kraglievich 1922 edentate Megalonychops primigenius Kraglievich 1926 edentate Torcellia paranense Kraglievich 1923 edentate "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum Bordas 1941 edentate Holotype: MACN Pv 13656, type of Ortotherium brevirostrum Bordas, 1942
Megatherioidea - Nothrotheriidae	Neohapalops rothi, Pronothrotherium mirabilis Neohapalops rothi Kraglievich 1923 edentate Pronothrotherium mirabilis Kraglievich 1925 edentate
Megatherioidea - Megatheriidae	Promegatherium smaltatum, Promegatherium parvulum, Pyramiodontherium sp.⁵, "Promegatherium remulsum" = Pyramiodontherium remulsum⁴, "Pliomegatherium lelongi" = Pliomegatherium lelongi, "Pliomegatherium paranensis" = Pliomegatherium lelongi⁴, Megatherium antiquum⁴, Eomegatherium nanum⁵ Promegatherium smaltatum Ameghino 1883 edentate Promegatherium parvulum Ameghino 1891 edentate Pyramiodontherium sp.⁵ Rovereto 1914 edentate "Promegatherium remulsum" = Pyramiodontherium remulsum⁴ Ameghino 1886 edentate "Pliomegatherium lelongi" = Pliomegatherium lelongi Kraglievich 1930 edentate "Pliomegatherium paranensis" = Pliomegatherium lelongi⁴ Kraglievich 1930 edentate Megatherium antiquum⁴ Ameghino 1885 edentate Eomegatherium nanum⁵ Burmeister 1891 edentate
- Mylodontidae	Octomylodon aversus, "Sphenotherus paranensis" = Sphenotherus zavaletianus, Scelidotheriinae indet., Prolestodon paranensis, Prolestodon antiquus, Strabosodon acuticavus, Strabosodon obtusicavus, Promylodon paranensis, Megabradys darwini, Ranculcus scalabrinianus Octomylodon aversus Ameghino 1904 edentate "Sphenotherus paranensis" = Sphenotherus zavaletianus Ameghino 1891 edentate Scelidotheriinae indet. Ameghino 1889 edentate Prolestodon paranensis Ameghino 1889 edentate Prolestodon antiquus Ameghino 1885 edentate Strabosodon acuticavus Ameghino 1891 edentate Strabosodon obtusicavus Ameghino 1891 edentate Promylodon paranensis Ameghino 1883 edentate Megabradys darwini Scillato-Yané 1981 edentate Ranculcus scalabrinianus Ameghino 1891 edentate
Notoungulata - Interatheriidae	Protypotherium antiquum, Munizia paranensis Protypotherium antiquum Ameghino 1882 notoungulate Munizia paranensis Kraglievich 1931 notoungulate
Notoungulata - Mesotheriidae	Eutypotherium sp.²¹ Eutypotherium sp.²¹ Roth 1902 notoungulate
Notoungulata - Haplodontheriidae	"Haplodontherium wildei" = Haplodontherium wildei, "Toxodontherium compressum" = Haplodontherium wildei, "Haplodontherium limun" = Haplodontherium wildei, "Toxodontherium reverendum" = Haplodontherium wildei, Pachynodon modicus "Haplodontherium wildei" = Haplodontherium wildei Ameghino 1885 notoungulate "Toxodontherium compressum" = Haplodontherium wildei Ameghino 1885 notoungulate "Haplodontherium limun" = Haplodontherium wildei Ameghino 1885 notoungulate "Toxodontherium reverendum" = Haplodontherium wildei Ameghino 1885 notoungulate Pachynodon modicus Burmeister 1891 notoungulate
Notoungulata - Toxodontidae	? Palaeotoxodon protoburmeisteri, "Stenotephanos plicidens" = Stenotephanos plicidens, "Eutomodus elautus" = Stenotephanos plicidens, "? Palaeotoxodon virgatus" = Stenotephanos plicidens, "Dinotoxodon paranensis" = Dinotoxodon paranensis, "Palaeotoxodon paranensis" = Dinotoxodon paranensis, ? Adinotherium paranense, ? Berroia sp., Dilobodon lutarius, Xotodon doellojuradi, Xotodon foricurvatus ? Palaeotoxodon protoburmeisteri Ameghino 1887 notoungulate "Stenotephanos plicidens" = Stenotephanos plicidens Ameghino 1885 notoungulate "Eutomodus elautus" = Stenotephanos plicidens Ameghino 1885 notoungulate "? Palaeotoxodon virgatus" = Stenotephanos plicidens Ameghino 1885 notoungulate "Dinotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate "Palaeotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate ? Adinotherium paranense notoungulate ? Berroia sp. Kraglievich 1931 notoungulate Dilobodon lutarius Ameghino 1886 notoungulate Xotodon doellojuradi Frenguelli 1920 notoungulate Xotodon foricurvatus Ameghino 1885 notoungulate
Rodentia - Erethizontidae	Paradoxomys sp., Paradoxomys cancrivorus⁷ Paradoxomys sp. Ameghino 1885 porcupine Paradoxomys cancrivorus⁷ Ameghino 1885 porcupine
Rodentia - Chinchillidae	"Lagostomopsis sp." = Lagostomus (Lagostomopsis), "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)⁷, "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)⁷ "Lagostomopsis sp." = Lagostomus (Lagostomopsis) Kraglievich 1926 viscacha "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)⁷ viscacha "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)⁷ viscacha
Rodentia - Caviidae	Caviodon sp., Parodimys sp., Prodolichotis sp., Cardiomys sp., Palaeocavia sp., Pliodolichotis sp.⁶ Caviodon sp. Ameghino 1885 caviomorph Parodimys sp. Kraglievich 1932 caviomorph Prodolichotis sp. Kraglievich 1932 caviomorph Cardiomys sp. Ameghino 1885 caviomorph Palaeocavia sp. Ameghino 1889 caviomorph Pliodolichotis sp.⁶ Kraglievich 1927 mara
Rodentia - Hydrochoeridae	Contracavia sp., "Cardiatherium sp." = Cardiatherium, Cardiatherium paranense n. sp.¹⁸, "Kiyutherium sp." = Cardiatherium, "Procardiatherium sp." = Cardiatherium, "Plexochoerus sp." = Cardiatherium, "Anchimys sp." = Cardiatherium, Anatochoerus sp., Cantracavia sp.⁷, "Protohydrochoerus sp." = Phugatherium⁷, Phugatherium cataclisticum, "Anchimysops sp." = Phugatherium Contracavia sp. Burmeister 1885 caviomorph "Cardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph Cardiatherium paranense n. sp.¹⁸ Ameghino 1883 caviomorph "Kiyutherium sp." = Cardiatherium Ameghino 1883 caviomorph "Procardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph "Plexochoerus sp." = Cardiatherium Ameghino 1883 caviomorph "Anchimys sp." = Cardiatherium Ameghino 1883 caviomorph Anatochoerus sp. Vucetich and Mones 1991 caviomorph Cantracavia sp.⁷ Burmeister 1885 caviomorph "Protohydrochoerus sp." = Phugatherium⁷ Ameghino 1887 caviomorph Phugatherium cataclisticum Ameghino 1887 caviomorph "Anchimysops sp." = Phugatherium Ameghino 1887 caviomorph
Rodentia - Dinomyidae	Tetrastylomys castellanosi⁷, Pseudosigmomys sp., Pseudosigmomys paranensis⁷, Rusconia crassidens⁷, Olenopsis typicus⁷, Potamarchus sp., Potamarchus sigmodon⁷, Potamarchus murinus⁷, Briaromys sp., Briaromys trouessartianus⁷, Paranamys sp., "Drytomomys typicus" = Paranamys typicus⁸, Eumegamys sp., Eumegamys dubius⁷, Eumegamys depressidens⁷, Eumegamys paranensis⁷, Eumegamys contortus⁷, Eumegamys scalabrinianus⁷, Doellomys sp., Doellomys parcus⁷, Carlesia sp., Carlesia pendolai⁷, Pentastylomys sp., Pentastylomys seriei⁷, Telodontomys compressidens⁷, Colpostemma sp., Pentastylodon racedi⁷, Gyriabrus sp., Gyriabrus glutinatus⁷, Gyriabrus indivisus⁷, Gyriabrus rebagliattii⁷, Gyriabrus holmbergi⁷, Eumegamysops sp., Eumegamysops praependens⁷, Diaphoromys sp., Diaphoromys compressidens⁷, Diaphoromys mesopotamicus⁷, Diaphoromys gamayensis⁷, Tetrastylus aguilari⁷, Tetrastylus diffusus⁷, Tetrastylus robustus⁷, Tetrastylus atavus⁷, Tetrastylus laevigatus⁷, Tetrastylus laevigatus n. sp.²⁴, Isostylomys sp., Isostylomys ameghinoi⁷, Isostylomys laurillardi⁷, Discolomys cuneus⁷, Protomegamys sp., Protomegamys coligatus⁷ Tetrastylomys castellanosi⁷ caviomorph Pseudosigmomys sp. Kraglievich 1931 caviomorph Pseudosigmomys paranensis⁷ caviomorph Rusconia crassidens⁷ Kraglievich 1931 caviomorph Olenopsis typicus⁷ Ameghino 1889 caviomorph Potamarchus sp. Burmeister 1885 caviomorph Potamarchus sigmodon⁷ caviomorph Potamarchus murinus⁷ Burmeister 1885 caviomorph Briaromys sp. Ameghino 1889 caviomorph Briaromys trouessartianus⁷ Ameghino 1889 caviomorph Paranamys sp. Kraglievich 1934 caviomorph "Drytomomys typicus" = Paranamys typicus⁸ Ameghino 1889 caviomorph Eumegamys sp. Kraglievich 1926 caviomorph Eumegamys dubius⁷ caviomorph Eumegamys depressidens⁷ caviomorph "Eumegamys" depressidens Eumegamys paranensis⁷ Kraglievich 1926 caviomorph Eumegamys contortus⁷ Ameghino 1889 caviomorph Eumegamys scalabrinianus⁷ Kraglievich 1926 caviomorph Doellomys sp. Alvarez 1947 caviomorph Doellomys parcus⁷ caviomorph Carlesia sp. Kraglievich 1926 caviomorph Carlesia pendolai⁷ caviomorph Pentastylomys sp. Kraglievich 1926 caviomorph Pentastylomys seriei⁷ caviomorph Telodontomys compressidens⁷ Kraglievich 1931 caviomorph Colpostemma sp. Ameghino 1891 caviomorph Pentastylodon racedi⁷ caviomorph Gyriabrus sp. Ameghino 1891 caviomorph Gyriabrus glutinatus⁷ caviomorph Gyriabrus indivisus⁷ caviomorph Gyriabrus rebagliattii⁷ caviomorph Gyriabrus holmbergi⁷ Ameghino 1885 caviomorph Eumegamysops sp. Alvarez 1947 caviomorph Eumegamysops praependens⁷ Ameghino 1886 caviomorph Diaphoromys sp. Kraglievich 1931 caviomorph Diaphoromys compressidens⁷ caviomorph Diaphoromys mesopotamicus⁷ Kraglievich 1932 caviomorph Diaphoromys gamayensis⁷ Kraglievich 1931 caviomorph Tetrastylus aguilari⁷ caviomorph Tetrastylus diffusus⁷ caviomorph Tetrastylus robustus⁷ caviomorph Tetrastylus atavus⁷ caviomorph T. (Protelicomys) atavus Tetrastylus laevigatus⁷ Ameghino 1886 caviomorph Tetrastylus laevigatus n. sp.²⁴ Ameghino 1886 caviomorph Isostylomys sp. Kraglievich 1926 caviomorph Isostylomys ameghinoi⁷ caviomorph Isostylomys laurillardi⁷ Ameghino 1883 caviomorph Discolomys cuneus⁷ Ameghino 1889 caviomorph "Discolomys" Protomegamys sp. Kraglievich 1932 caviomorph Protomegamys coligatus⁷ caviomorph
Rodentia - Neoepiblemidae	Perimys scalabrinianus⁷, Neoepiblema sp., Epiblema horridula¹⁴, "Neoepiblema ambrosettianus" = Neoepiblema horridula¹⁹, Phoberomys sp., Phoberomys sp.⁷, "Phoberomys burmeisteri" = Phoberomys burmeisteri¹⁵, "Phoberomys insolita n. sp." = Phoberomys burmeisteri¹⁷, "Megamys burmeisteri" = Phoberomys burmeisteri¹⁴, "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri¹⁷, "Phoberomys minima n. sp." = Phoberomys burmeisteri¹⁷, "Phoberomys minima" = Phoberomys burmeisteri⁷, "Phoberomys praecursor n. sp." = Phoberomys burmeisteri¹⁶ Perimys scalabrinianus⁷ Ameghino 1889 caviomorph Neoepiblema sp. Ameghino 1889 caviomorph Epiblema horridula¹⁴ Ameghino 1886 caviomorph upper right molar - collected by F. Ameghino 1886 "Neoepiblema ambrosettianus" = Neoepiblema horridula¹⁹ Ameghino 1886 caviomorph Phoberomys sp. Kraglievich 1926 caviomorph Phoberomys sp.⁷ Kraglievich 1926 caviomorph "Phoberomys burmeisteri" = Phoberomys burmeisteri¹⁵ Kraglievich 1926 caviomorph "Phoberomys insolita n. sp." = Phoberomys burmeisteri¹⁷ Kraglievich 1926 caviomorph "Megamys burmeisteri" = Phoberomys burmeisteri¹⁴ Kraglievich 1926 caviomorph collected by F. Ameghino 1886 "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri¹⁷ Kraglievich 1926 caviomorph "Phoberomys minima n. sp." = Phoberomys burmeisteri¹⁷ Kraglievich 1926 caviomorph "Phoberomys minima" = Phoberomys burmeisteri⁷ Kraglievich 1926 caviomorph "Phoberomys praecursor n. sp." = Phoberomys burmeisteri¹⁶ Kraglievich 1926 caviomorph
Rodentia - Echimyidae	Haplostropha sp., Haplostropha sacabriniana⁷, Eumysops parodii, Eumysops parodii⁷, Myocastor sp., Myocastor paranensis⁷, Myocastor obesus⁷, Myocastor sinuata⁷ Haplostropha sp. Ameghino 1891 spiny rat Haplostropha sacabriniana⁷ spiny rat Eumysops parodii Kraglievich 1945 spiny rat Eumysops parodii⁷ Kraglievich 1945 spiny rat “Eumysops” parodii Myocastor sp. Kerr 1782 spiny rat Myocastor paranensis⁷ spiny rat Myocastor obesus⁷ spiny rat Myocastor sinuata⁷ spiny rat
Rodentia - Abrocomidae	Protabrocoma sp., Protabrocoma paranensis⁷ Protabrocoma sp. Kraglievich 1927 chinchilla rat Protabrocoma paranensis⁷ chinchilla rat
Rodentia - Myocastoridae	Strophostephanos sp.⁶, Strophostephanos jheringi⁷ Strophostephanos sp.⁶ Ameghino 1891 caviomorph Strophostephanos jheringi⁷ caviomorph
Carnivora - Procyonidae	Cyonasua argentina Cyonasua argentina Ameghino 1885 procyonid carnivore
Panameriungulata - Proterotheriidae	Neobrachytherium ameghinoi²¹, "Proterotherium gradatum" = Brachytherium gradatum, "Brachytherium cuspidatum" = Brachytherium cuspidatum, "Licaphrium mesopotamiense" = Brachytherium cuspidatum, "Epitherium paranensis" = Brachytherium cuspidatum, Proterotherium cervioides, "Coelosoma eversa" = Diadiaphorus eversus Neobrachytherium ameghinoi²¹ Soria 2001 placental "Proterotherium gradatum" = Brachytherium gradatum Ameghino 1891 placental "Brachytherium cuspidatum" = Brachytherium cuspidatum Ameghino 1883 placental "Licaphrium mesopotamiense" = Brachytherium cuspidatum Ameghino 1883 placental "Epitherium paranensis" = Brachytherium cuspidatum Ameghino 1883 placental Proterotherium cervioides Ameghino 1883 placental "Coelosoma eversa" = Diadiaphorus eversus Soria 2001 placental
Panameriungulata - Macraucheniidae	Paranauchenia denticulata, Promacrauchenia antiqua, Cullinia sp.²¹, "Oxyodontherium zeballosi" = Oxyodontherium zeballozi, "Mesorhinus piramydatus" = Oxyodontherium piramydatus, Scalabrinitherium bravardi, Scalabrinitherium rothii Paranauchenia denticulata Ameghino 1891 placental Promacrauchenia antiqua Ameghino 1887 placental Cullinia sp.²¹ Cabrera and Kraglievich 1931 placental "Oxyodontherium zeballosi" = Oxyodontherium zeballozi Ameghino 1883 placental "Mesorhinus piramydatus" = Oxyodontherium piramydatus Ameghino 1885 placental Scalabrinitherium bravardi Ameghino 1883 placental Scalabrinitherium rothii Ameghino 1885 placental
Cetacea - Iniidae	Iniidae indet.¹³, "Saurodelphis gigas n. sp." = Saurocetes gigas⁹ Iniidae indet.¹³ Gray 1846 toothed whale "Saurodelphis gigas n. sp." = Saurocetes gigas⁹ Cozzuol 1988 toothed whale
Sparassodonta -	Stylocynus paranensis, "Notictis ortizi" = Notictis ortizii Stylocynus paranensis Mercerat 1917 metatherian "Notictis ortizi" = Notictis ortizii Ameghino 1889 metatherian
Sparassodonta - Thylacosmilidae	Thylacosmilus atrox¹² Thylacosmilus atrox¹² Riggs 1933 metatherian
Didelphimorphia - Didelphidae	Philander entrerianus¹², Chironectes sp.¹², Zygolestes paranensis¹² Philander entrerianus¹² Ameghino 1899 four-eyed opossum Chironectes sp.¹² Illiger 1811 water opossum Zygolestes paranensis¹² Ameghino 1898 opossum
Actinopteri
Characiformes - Cynodontidae	Cynodontidae indet. Cynodontidae indet.
Characiformes - Characidae	Colossoma macropomus Colossoma macropomus

¹Bona and Barrios 2015; ²Bona et al. 2024; ³Bona and Carabajal 2013; ⁴Brandoni 2006; ⁵Brandoni 2013; ⁶Brandoni 2013; ⁷Candela 2005; ⁸Candela and Nasif 2006; ⁹Cozzuol 1988; ¹⁰Falbo and Agnolin 2020; ¹¹Gasparini 1985; ¹²Goin et al. 2013; ¹³Gutstein et al. 2014; ¹⁴Kerber et al. 2019; ¹⁵Kraglievich 1926; ¹⁶Kraglievich 1932; ¹⁷Kraglievich 1940; ¹⁸Mones 1991; ¹⁹Negri and Ferigolo 1999; ²⁰Nuñez-Blasco et al. 2023; ²¹Schmidt and Cerdeño 2013; ²²Scillato-Yané 1975; ²³Scillato-Yané et al. 2013; ²⁴Sostillo et al. 2022