SH-20, Khodzhakul (Cretaceous of Uzbekistan)

Also known as Khodja-Kull; SKH-20

Where: Navoi, Uzbekistan (40.1° N, 65.8° E: paleocoordinates 35.9° N, 60.5° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Khodzhakul Formation, Early/Lower Cenomanian (99.6 - 93.5 Ma)

• lower/middle part of the formation

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; pebbly, calcareous sandstone and pebbly, sandy claystone

• "limonite pebbles lying amongst a suite of calcareous sandstones and sandy clays"

Size classes: macrofossils, mesofossils

Collected by A. D. Arkhangelsky, L. A. Nessov in 1914; 1978, 1981-1982

Primary reference: A. N. Riabinin. 1931. Ostatki dinozavrov iz verkhnego mela nizov'ev r. Amu-Dar'i [On the dinosaurian remains from the Upper Cretaceous of the lower parts of the Amu-Daria River]. Zapiski Rossiyskogo Mineralogicheskogo Obshchestva 60(1):114-118 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 58208: authorized by Matthew Carrano, entered by Kaitlin Maguire on 27.01.2006, edited by Roger Benson, Matthew Carrano, John Alroy and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Malacostraca indet.12 Latreille 1802
Bivalvia indet.12 Linnaeus 1758 clam
Gastropoda indet.12 Cuvier 1797 snail
 Ammonoidea -
 Hybodontiformes - Polyacrodontidae
 Hybodontiformes - Distobatidae
Glickmanodus rarus n. gen. n. sp.9
Glickmanodus rarus n. gen. n. sp.9 Nessov et al. 1997 elasmobranch
including holotype: CCMGE 677/12457
 Euselachii -
 Lamniformes - Anacoracidae
"Eoanacorax dalinkevichiusi" = Squalicorax dalinkevichiusi12
"Eoanacorax dalinkevichiusi" = Squalicorax dalinkevichiusi12 Glikman and Shvazhaite 1971 crow shark
 Lamniformes - Cretoxyrhinidae
Paraisurus sp.12 Glikman 1957 mackerel shark
 Holocephali -
 Acipenseriformes - Polyodontidae
Polyodontidae indet.9 Bonaparte 1838 paddlefish
Hesperopsephurus kyatensis n. gen. n. sp.10 Nessov 1997 paddlefish
including holotype CCMGE 32/12177
 Teleostei - Aspidorhynchidae
 Salmoniformes - Enchodontidae
Enchodontidae indet.7 Woodward 1901
Eurypholis sp.9 Pictet 1850
 Salientia -
 Caudata - Urodela
Horezmia gracile4 Nessov 1981 salamander
 Testudines - Macrobaenidae
Oxemys gutta n. gen. n. sp.5, cf. Kirgizemys sp.5
Oxemys gutta n. gen. n. sp.5 Nessov 1977 turtle
TsNIGR Museum, no. 2/11478
cf. Kirgizemys sp.5 Nessov and Khozatsky 1973 turtle
 Testudines -
"Chelonia indet." = Testudines12
"Chelonia indet." = Testudines12 Batsch 1788 turtle
 Testudines - Nanhsiungchelyidae
Nanhsiungchelyidae indet.2 Yeh 1966 turtle
 Testudines - Carettochelyidae
Kizylkumemys sp.5 Nessov 1977 turtle
 Testudines - Lindholmemydidae
"Mongolemys cf. occidentalis" = Khodzhakulemys occidentalis5
"Mongolemys cf. occidentalis" = Khodzhakulemys occidentalis5 Nessov 1984 turtle
 Testudines - Adocidae
Adocus kizylkumensis4 Nesov 1981 turtle
 Saurischia -
Neosauropoda indet.9 Bonaparte 1986 sauropod
"Diplodocidae or Titanosauridae"
Aves indet.12, Horezmavis eocretacea n. gen. n. sp.11
Aves indet.12 Linnaeus 1758 bird
Horezmavis eocretacea n. gen. n. sp.11 Nessov and Borkin 1983 bird
 Avetheropoda -
"Laelaps" cf. explanatus9 Cope 1876 coelurosaur
"possibly this is a peculiar species of Alectrosaurus"
 Avetheropoda - Coeluridae
Coeluridae indet.9 Marsh 1881 coelurid
 Ornithischia - Ankylosauridae
? Shamosaurinae indet.9 Tumanova 1983 ornithischian
also listed as Shamosauridae
 Neornithischia -
Kulceratops kulensis n. gen. n. sp.9
Kulceratops kulensis n. gen. n. sp.9 Nessov 1995 ornithischian
CCMGE 495/12457
 Neornithischia - Hadrosauridae
Hadrosauridae indet. Cope 1869 hadrosaurid
 Neornithischia - Hypsilophodontidae
? Hypsilophodontidae indet.9 Dollo 1882 hypsilophodontid
 Neornithischia -
? Gilmoreosaurus atavus n. sp.9 Nessov 1995 bird foot dinosaur
CCMGE 576/12457
 Loricata -
Crocodylia indet.12 crocodilian
 Squamata -
Iguania indet.3 Cuvier 1817 lizard
CMGE 40/12000 (frontal), 41/12000 (partial parietal)
Gekkota indet.9 Cuvier 1817 lizard
Scincomorpha indet.3 Camp 1923 lizard
CMGE 42/12000 (partial right dentary)
Oxia karakalpakensis n. gen. n. sp.7 Nessov 1985 lizard
Dentaries: CMGE 28/12000 (holotype; Nessov 1985); 29/12000, 43/12000, 44/12000
Hodzhakulia magna n. gen. n. sp.7 Nessov 1985 worm lizard
Maxillae: CMGE30/12000 (holotype, right), 31/12000 (left); right dentaries: 32/12000, 32a/12000
 Tribosphenida -
Bobolestes zenge n. gen. n. sp.6
Bobolestes zenge n. gen. n. sp.6 Nessov 1985 placental
from "Khodzhakul, SKH-20"; CCMGE 2/12176
 Tribosphenida - Zhelestidae
cf. Sheikhdzheilia rezvyii1 Averianov and Archibald 2005 eutherian
from "Khodzhakul, SKH-20"
 Theriiformes - Pappotheriidae
Pappotheriidae sp.8 Slaughter 1965 metatherian
 Pinales - Araucariaceae
Araucariaceae indet.9 Henckel and Hochstetter 1865
 Pinales -
"Coniferales indet." = Pinales12
"Coniferales indet." = Pinales12 Dumortier 1829
Angiospermae indet.12 Mabberley 2000