SH-5, Khodzhakulsai ravine (Cretaceous of Uzbekistan)

Also known as Khodzhakulsay, SKh-5

Where: Karakalpakstan, Uzbekistan (40.1° N, 65.8° E: paleocoordinates 35.9° N, 60.5° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Khodzhakul Formation, Early/Lower Cenomanian (99.6 - 93.5 Ma)

• Upper part of the Khodzhakul formation

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone

Size classes: macrofossils, mesofossils

Primary reference: L. A. Nessov. 1984. Data on late Mesozoic turtles from the USSR. Studia Geologica Salamanticensia, vol. especial 1 (Studia Palaeocheloniologica I) 1:215-223 [R. Benson/R. Benson]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 58490: authorized by Matthew Carrano, entered by Kaitlin Maguire on 09.02.2006, edited by Richard Butler, Matthew Carrano, Roger Benson and Jonathan Tennant

Creative Commons license: CC BY (attribution)

Taxonomic list

• Nessov (1984) listed turtles from "Sheichdzheili and Chodzhakul localities and others", but only gave details for SH-5
Actinopteri
 Albuliformes - Albulidae
Cretalbula hammada4 Nessov 1985 bonefish
 Pycnodontiformes - Pycnodontidae
Anomoeodus splendidus4 Nessov 1985
 Amiiformes - Amiidae
Amia semimarina4 Nessov 1985 bowfin
Reptilia
 Testudines - Macrobaenidae
Anatolemys oxensis Khozatskii and Nessov 1977 turtle
cf. Kirgizemys sp. Nessov and Khozatsky 1973 turtle
 Testudines - Trionychidae
Trionyx "species 3" Geoffroy Saint-Hilaire 1809 softshell turtle
Trionyx dissolutus6 Vitek and Danilov 2014 softshell turtle
 Testudines -
Tienfucheloides undatus Nessov 1978 turtle
 Testudines - Adocidae
Adocus kizylkumensis2 Nesov 1981 turtle
"Ferganemys" itemirensis2 Nesov 1981 turtle
Pie chart in Nessov (1984) suggests ~250 specimens
 Testudines - Carettochelyidae
Kizylkumemys schultzi n. gen. n. sp.
Kizylkumemys schultzi n. gen. n. sp. Nessov 1977 turtle
 Testudines - Pantrionychidae
"Trionyx cf. kyrgyzensis" = Petrochelys kyrgyzensis6
"Trionyx cf. kyrgyzensis" = Petrochelys kyrgyzensis6 Nessov 1995 turtle
 Testudines - Lindholmemydidae
"Mongolemys occidentalis" = Khodzhakulemys occidentalis
"Mongolemys occidentalis" = Khodzhakulemys occidentalis Nessov 1984 turtle
 Avetheropoda -
? Deinonychosauria indet.5 Colbert and Russell 1969 coelurosaur
Aves
 Alexornithiformes - Alexornithidae
Zhyraornis kashkarovi3 Nessov 1984 bird
TsNIGRI 43/11915, thoracic vertebra
Reptilia
 Avetheropoda -
Alectrosaurus sp.5 Gilmore 1933 coelurosaur
 Neornithischia -
Asiaceratops salsopaludalis5 Nessov et al. 1989 ornithischian
 Neornithischia - Hadrosauridae
Hadrosauridae indet.5 Cope 1869 hadrosaurid
 Pterosauria - Ornithocheiridae
Ornithocheiridae indet.1 pterosaur
ZIN PH 3/44, tooth