PPP# 170 - Escudo de Veraguas [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 17622; 87-09-03

Where: Bocas, Panama (9.1° N, 81.5° W: paleocoordinates 9.8° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: NN15 nannofossil zone, Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), planktic foraminifera suggest 1.8 Ma near the top and 3.5 Ma at the base. PPP dataset indicates an age range between 3.6 and 3.5 Ma, nannofossil zone NN15.

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, prevasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 8/7/87. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 61337: authorized by Austin Hendy, entered by Austin Hendy on 05.06.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
unclassified
  -
? Turbonillidae indet.
originally entered as "? Turbonillidae sp."
Scaphopoda
 Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758 tusk shell
sensu-lato
 Gadilida - Entalinidae
Bathoxiphus sp. Pilsbry 1897 tusk shell
 Gadilida - Gadilidae
Gadilopsis sp. Woodring 1925 tusk shell
Gastropoda
 Heterostropha - Pyramidellidae
Pyramidellidae indet. Gray 1840 snail
Eulimella (Ebalina) sp. Thiele 1929 snail
 Heterostropha - Architectonicidae
Architectonica sp. Röding 1798 sundial
Pseudotorinia sp. Sacco 1892 snail
 Heterostropha - Acteonidae
Acteon sp. Montfort 1810 snail
 Trochoidea - Trochidae
Microgaza sp. Dall 1881 top snail
 Thecosomata - Cavoliniidae
Cavolinia sp. Avilgaard 1791 snail
Styliola sp. Gray 1850 snail
 Opisthobranchia - Retusidae
Volvulella sp. Newton 1891 snail
 Opisthobranchia - Cylichnidae
Acteocina sp. Gray 1847 barrel-bubble
 Cocculinoidea - Cocculinidae
? Cocculina sp. Dall 1882 snail
 Sorbeoconcha - Naticidae
"Natica (Naticarius) sp." = Naticarius
"Natica (Naticarius) sp." = Naticarius Dumeril 1806 moon snail
 Sorbeoconcha - Rissoidae
Rissoidae indet. Gray 1847 snail
 Neogastropoda - Columbellidae
Cotonopsis sp. Olsson 1942 snail
 Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806 snail
sensu-lato
 Neogastropoda - Buccinidae
Metula sp. Adams and Adams 1853 true whelk
Antillophos (Antillophos) sp. Woodring 1928 true whelk
 Neogastropoda - Drilliidae
Agladrillia (Agladrillia) sp. Woodring 1928 snail
 Neogastropoda - Mangeliidae
Ithycythara sp. Woodring 1928 snail
 Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891 auger snail
 Neogastropoda - Conidae
Conus sp. Linnaeus 1758 cone shell
 Neogastropoda - Turridae
Polystira sp. Woodring 1928 turrid
 Neogastropoda - Olividae
Olivella (Minioliva) sp. Olsson 1956 olive snail
Olivella (Macgintiella) sp. Olsson 1956 olive snail
Oliva (Oliva) sp. Bruguière 1789 olive snail
 Neogastropoda - Marginellidae
Marginellidae indet. Fleming 1828 margin shell
Volvarina sp. Hinds 1844 margin shell
 Neogastropoda - Muricidae
Haustellum sp. Schumacher 1817 murex snail
 Sorbeoconcha - Tonnidae
Malea ringens Swainson 1822 tun shell
 Sorbeoconcha - Personidae
Distorsio clathrata Lamarck 1816 snail
Distorsio decussata Valenciennes 1832 snail
 Sorbeoconcha - Bursidae
Marsupina bufo Bruguiére 1792 frog shell
 Sorbeoconcha - Cassidae
Sconsia grayi Adams 1855 snail
 Pseudomelanoidea - Pseudomelaniidae
Miraclathurella sp. Woodring 1928 snail
? Compsodrillia sp. Woodring 1928 snail
 Vanikoroidea - Eulimidae
Eulimidae indet. Philippi 1853 snail
Bivalvia
 Nuculanida - Nuculanidae
Saccella sp. Woodring 1925 pointed nut clam
 Nuculida - Nuculidae
Nucula (Lamellinucula) sp. Schenck 1944 nut clam
 Arcida - Arcidae
Anadara (Rasia) sp. Gray 1857 ark
Anthozoa
 Scleractinia - Caryophylliidae
Asterosmilia profunda Duncan 1864 stony coral