Berezovsk Quarry (Jurassic of Russian Federation)

Also known as Stegosaur Quarry; Berezovsk coal mine

Where: Krasnoyarsk, Russian Federation (55.7° N, 89.2° E: paleocoordinates 58.2° N, 86.7° E)

• coordinate stated in text

• small collection-level geographic resolution

When: Upper Member (Itat Formation), Bathonian (168.3 - 166.1 Ma)

• upper part of formation

•The age of the Upper Mem- ber of the Itat Formation is Bathonian according to the pollen complex, which consists of Cyathidites minor, Piceapollenites sp., Eboracia torosa, Quadraeculina limbata, and Classopollis sp. (Raevskaya et al. 1993)

• group of beds-level stratigraphic resolution

Environment/lithology: fluvial; gray claystone

• By mid-Jurassic times, a low-profile alluvial plain had developed in this area that was characterized by anas- tomosing rivers and well-defined floodplains (Le Heron et al. 2008). Due to the fluvial origin of the sediments, vertebrate remains are mostly disarticulated and often water worn. However, the predominantly good preserva- tion of vertebrate remains demonstrates that transport occurred over short distances.
• "grey clays"

•Fossil rich layer tops a thick coal seam

Size classes: macrofossils, mesofossils, microfossils

Collected by S. Krasnolutskii et al. in 2000-2005, 2010–2012; reposited in the PIN

Collection methods: bulk, quarrying, surface (in situ), mechanical, sieve,

• Itatodon tatarinovi material housed in the Paleontological Museum of Tomsk State

•University, Tomsk, Russia (PM TGU), and in the Paleontological Institute of the Russian Academy of Sciences, Moscow, Russia (PIN)

•2010-2012 collection by a Joint Russian-German expedition: "The newly collected mammal specimens were obtained during screen-washing of ~12.5 tons of matrix. The jaw fragments were found in the field by picking the coarse fraction of fossiliferous concentrate. The teeth were recovered during sorting of the fine fraction in the laboratory.

•The specimens described here are housed in the Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow (PIN).

•"...other specimens were collected in 2010–2012 during a joint Russian-German expedition to Berezovsk Quarry. The edentulous jaw fragments were found in the field while picking the coarse fraction of fossiliferous concentrate obtained from screenwashing of ∼12.5 tons of matrix" (Avaerianov et al. 2014)"

Primary reference: A. O. Averianov, A. V. Lopatin, P. P. Skutschas, N. V. Matynovich, S. V. Leschinkskiy, A. S. Rezvyi, S. A. Krasnolutskii and A. V. Fayngertz. 2005. Discovery of Middle Jurassic mammals from Siberia. Acta Palaeontologica Polonica 50(4):789-797 [P. Mannion/J. Tennant]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 61891: authorized by Matthew Carrano, entered by Matthew Carrano on 06.07.2006, edited by Oliver Rauhut, Roger Benson, Jonathan Tennant, Philip Mannion, Evangelos Vlachos and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

Chondrichthyes
 Hybodontiformes - Hybodontidae
Hybodus sp.8 Agassiz 1834 elasmobranch
Osteichthyes
 Palaeonisciformes -
 Dipnoi -
Ferganoceratodus sp.8 Nessov and Kaznyshkin 1985 lungfish
Reptilia
 Choristodera -
? Choristodera indet.11 choristodere
 Loricata - Goniopholididae
Goniopholididae indet.10 Cope 1875 crocodilian
 Pterosauria - Rhamphorhynchidae
Rhamphorhynchidae indet.8 Seeley 1870 pterosaur
 Ornithischia - Heterodontosauridae
Heterodontosauridae indet.14 Romer 1966 heterodontosaurid
 Ornithischia -
Stegosauria indet.14 ornithischian
ZIN PH 1–2/117; SHRM various
 Saurischia - Mamenchisauridae
Mamenchisauridae indet.8 mamenchisaurid
 Theropoda - Proceratosauridae
Kileskus aristotocus n. gen. n. sp.1 Averianov et al. 2010 coelurosaur
Kileskus aristotocus8 Averianov et al. 2010 coelurosaur
 Testudines -
Testudines "indet. 1"8 Batsch 1788 turtle
ZIN PH, no. 1/186, left costal VIII.
Testudines "indet. 2"9 Batsch 1788 turtle
ZIN PH, no. 2/186, left xiphiplastron fragment; and specimen ZIN PH, no. 3/186, shell fragment.
Testudines "indet. 3"9 Batsch 1788 turtle
ZIN PH, no. 4/186, distal part of a right humerus.
 Testudines - Xinjiangchelyidae
Annemys sp.8 Sukhanov and Narmandakh 2006 turtle
Annemys variabilis n. sp.13 Obraztsova et al. 2022 turtle
ZIN PH, no. 431/50, costal VIII; ZIN PH, nos. 1/207 and 2/207, medial parts of costal VIII; ZIN PH, nos. 14/187 and 17/187, xiphiplastra; ZIN PH, no. 3/207, xiphiplastron fragment; ZIN PH, no. 6/185, right humerus; and ZIN PH, no. 4/207, distal part of the left humerus.
 Squamata -
Scincomorpha indet.8 Camp 1923 squamates
Mammalia
  -
Mammalia indet. Linnaeus 1758 mammal
 Eutriconodonta -
Eutriconodonta indet.2 Kermack et al. 1973 mammal
PIN 5087/4, left upper molariform tooth
 Dryolestoidea - Dryolestidae
Anthracolestes sergeii n. gen. n. sp.5
Anthracolestes sergeii n. gen. n. sp.5 Averianov et al. 2014 mammal
 Cladotheria - Amphitheriidae
Amphibetulimus krasnolutskii n. gen. n. sp.12 Lopatin and Averianov 2007 mammal
PIN, no. 5087/3, right dentary fragment with M3 and alveoli and roots of M1, M2, M4, and M5; PIN 5087/34, right upper molar. PIN 5087/39, left dentary fragment with p1, alveoli or roots for i1–4, double rooted canine, and p2. PIN 5087/44, right dentary fragment with alveoli for two middle molars. PIN 5087/43, right dentary fragment with alveoli for ultimate and penultimate molars. PIN 5087/40, right dentary fragment with ultimate molar and roots of the penultimate molar. PIN 5087/45, right dentary fragment with alveolus for ultimate molar
 Multituberculata -
Tashtykia primaeva n. gen. n. sp.7, Tagaria antiqua n. gen. n. sp.7
Tashtykia primaeva n. gen. n. sp.7 Averianov et al. 2020 multituberculate
Holotype: PIN 5087/52, left p4; PIN 5087/25, right I2; LMCCE 010/2, right upper deciduous premolar; PIN 5087/46, right upper anterior premolar; PIN 5087/21, left P5
Tagaria antiqua n. gen. n. sp.7 Averianov et al. 2020 multituberculate
Holotype: PIN 5087/108, right P5; LMCCE 004/10, left M2; PIN 5087/50, right M2
 Allotheria -
Sineleutherus issedonicus n. sp.3 Averianov et al. 2011 mammal
PIN 5087/9 to 5087/18
Osteichthyes
 Docodonta - Tegotheriidae
Hutegotherium yaomingi n. gen. n. sp.4
Hutegotherium yaomingi n. gen. n. sp.4 Averianov et al. 2010 dodocont
PM TGU 200/5-BR-2; PIN 5087/6, 5087/8
 Docodonta - Simpsonodontidae
Simpsonodon sibiricus n. sp.4 Averianov et al. 2010 dodocont
PIN 5087/1, posterior fragment of a right denta− ry with alveoli for the ultimate molar, posterior alveolus for the penultimate molar and partially preserved coronoid and pseudangular processes. PM TGU 200/3−BR−1, posterior fragment of a right dentary with alveoli for the two last mo− lars and partially preserved coronoid process
 Docodonta -
Itatodon tatarinovi n. gen. n. sp.11
Itatodon tatarinovi n. gen. n. sp.11 Lopatin and Averianov 2005 dodocont
PIN 5087/2 (holotype, isolated right lower molar), 5087/7, PM TGU 200/3-BR-7
 Therapsida - Tritylodontidae
Stereognathus sp.6 Charlesworth 1854 mammaliamorph
Amphibia
 Caudata -
Caudata "indet. A"8 Scopoli 1777 salamander
Berezovsk salamander A
Egoria malashichevi n. gen. n. sp.15 Skutschas et al. 2020 salamander
 Caudata - Urodela
Kiyatriton krasnolutskii n. sp.8 Skutchas 2016 salamander
 Caudata - Karauridae
cf. Karauridae indet.14 Ivachnenko 1978 salamander
 Caudata -
Urupia monstrosa n. gen. n. sp.16 Skutschas and Krasnolutskii 2011 salamander
ZIN PH 1/144 (holotype atlas), 3/144, 4/144 (dorsal vertebrae); PM TGU 200.10, ZIN PH 2/144 (anterior dentary fragments); PM TGU 200/11 (femur)
Urupia monstrosa8 Skutschas and Krasnolutskii 2011 salamander
 Salientia - Discoglossidae
? Eodiscoglossus sp.17 Villalta 1957 painted frog
ZIN PH 27/144
Osteichthyes
 Halecomorphi - Sinamiidae
Sinamiidae indet.8 Berg 1940 ray-finned fish
 Chondrostei -
Acipenseriformes indet.8 Berg 1940 ray-finned fish