Bore Lullingen 1, depth 33m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61903: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Foraminifera
 Rotalioidea - Elphidiidae
Elphidium inflatum
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides pseudoungerianus, Cibicides dutemplei
Cibicides lobatulus
incl. 1 specimen C. lob. var. ornatus
Cibicides dutemplei dOrbigny 1846
 Rotaliida - Gavelinellidae
Gyroidina soldanii
Tubothalamea
 Miliolida - Miliolidae
Sigmoilina tenuis
Foraminifera
  - Spiroplectamminidae
Spiroplectammina carinata
Nodosariata
 Vaginulinida - Vaginulinidae
Lenticulina (Lenticulina) calcar, Lenticulina (Planularia) grundensis, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) orbicularis, Lenticulina (Robulus) cassis
 Nodosariida - Nodosariidae
Vaginulina legumen, Dentalina konincki, Nodosaria bacillum
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Globulina gibba, Guttulina problema, Pseudopolymorphina subnodosa
Globulina gibba d'Orbigny 1832
Guttulina problema d'Orbigny 1826
Foraminifera
 Lituolida - Lituolidae
Textularia gramen
Textularia gramen dOrbigny 1846
Scaphopoda
 Gadilida - Gadilidae
Gadila dingdenensis1
 Dentaliida - Dentaliidae
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Lentigodentalium pseudomutabile1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
Cephalopoda
 Spirulida - Spirulirostridae
Spirulirostra hoernesi1
Gastropoda
 Epitonioidea - Epitoniidae
Opalia (Pliciscala) pertusa2, Opalia (Nodiscala) rugatina2, "Amaea (Scalina) subreticula" = Scalina2, Epitonium (Spiniscala) frondiculum2, sensu lato Cirsotrema "sp. 1"2, Cirsotrema (Cirsotrema) crassicostatum2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
sensu lato Cirsotrema "sp. 1"2 Mörch 1852 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Vanikoroidea - Eulimidae
Niso (Niso) terebellum2, Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
 Cerithioidea - Cerithiidae
Bittium (Bittium) tenuispina2, "Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Sorbeoconcha - Naticidae
Naticidae indet.2, "Natica neglecta" = Cochlis neglecta2, "Natica sensu lato tigrina" = Cochlis tigrina2, Euspira edithae2, Euspira gottschei2, Euspira staringi2, Euspira helicina2, Polinices (Polinices) miocolligens2, Neverita josephinia2, Tectonatica miopusilla2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
Euspira edithae2 moon snail
Euspira gottschei2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Rissoidae
Alvania (Alvania) partschi2, Alvania (Alvania) basisulcata2, Ovirissoa gottscheana2
 Sorbeoconcha - Cassidae
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2, "Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) aperta2, Trigonostoma (Trigonostoma) extractrix2, Babylonella fusiformis2, Sveltia varicosa2, Cancellaria (Merica) contorta2, Cancellaria (Bivetiella) cancellata2
Sveltia varicosa2 snail
S. varicosa paucicostata
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Cancellaria (Bivetiella) cancellata2 Linnaeus 1767 snail
C. cancellata praecedens
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Columbellidae
Mitrella (Macrurella) nassoides2, Anachis (Costoanachis) hosiusi2
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Muricidae
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, Pterynotus (Alipurpura) parvifolius2, Coralliophila (Aldrichia) cf. granifera2, Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3, Terebra (Myurellina) acuminata3
Strioterebrum ronaldi3 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Mangeliidae
Benthomangelia aff. obtusangula3, Sorgenfreispira tenella3
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Raphitomidae
Teretia anceps3, Mioawateria andersoni3, Favriella sinuosula3, Metuonella grippi3, Pleurotomella "sp. 1"3
Pleurotomella "sp. 1"3 Verrill 1872 snail
 Neogastropoda - Turridae
Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula hanseata3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) sp.3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3, Ingaunoturricula marginata3
Gemmula (Gemmula) sp.3 Weinkauff 1875 turrid
Gemmula n. sp.
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Borsoniidae
Borsonia uniplicata3, "Genota ramosa" = Genota valeriae3, Asthenotoma pannoides3, Asthenotoma pannus3, Asthenotoma festiva3, Bathytoma jugleri3
Borsonia uniplicata3 Koenen 1872 snail
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
12 specimen C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Tornidae
Tornus (Tornus) trigonostoma2
 Heterostropha - Mathildidae
Gegania cancellata3, sensu lato Mathilda concinna3
 Heterostropha - Pyramidellidae
Chrysallida (Pyrgulina) pygmaea3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Turbonilla (Turbonilla) "sp. 2"3, Turbonilla (Mormula) "sp. 3"3, Turbonilla (Mormula) "sp. 4"3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) cf. pallidaeformis3, Odostomia (Megastomia) tuexeni3, Syrnola dubia3, Syrnola hoernesi3, Syrnola neumayri3, Syrnola "sp. 1"3
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros3 Sacco 1892 snail
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
Turbonilla (Mormula) "sp. 3"3 Adams 1864 snail
Turbonilla (Mormula) "sp. 4"3 Adams 1864 snail
Syrnola "sp. 1"3 Adams 1860 snail
 Heterostropha - Acteonidae
Acteon (Acteon) laevigatus3, Acteon (Acteon) semistriatus3, Acteon (Acteon) arnumensis3
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) arnumensis3 Sorgenfrei 1958 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3
 Opisthobranchia - Cylichnidae
Bullichna paucistriata3, Cylichna (Cylichna) pseudoconvoluta3, Scaphander grateloupi3
 Opisthobranchia - Bullidae
"Tornatina bellardi" = Bulla (Tornatina)3
"Tornatina bellardi" = Bulla (Tornatina)3 snail
 Opisthobranchia -
Roxania utriculus3
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1, Nuculana (Jupiteria) curvirostris1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
Nuculana (Jupiteria) curvirostris1 de Cserna 1961 pointed nut clam
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
incl. 1specimen of Portlandia pygm. fa. alta
 Nuculida - Nuculidae
Nuculoma haesendoncki1, Nucula (Nucula) nucleus1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Solenida - Pharidae
Cultellus (Cultellus) aff. cultellatus1
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, Callista chione1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Callista chione1 Linnaeus 1758 venus clam
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Lutraria frangula1
 Cardiida - Erycinidae
Erycina aff. backlundi1
 Cardiida - Montacutidae
Montacuta cf. caecilia1
 Cardiida - Solecurtidae
Solecurtus basteroti1, Azorinus (Azorinus) chamasolen1
Azorinus (Azorinus) chamasolen1 clam
A. chamasolen miocaenicus
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Semelidae
Abra (Abra) antwerpiensis1, Abra (? Abra) sorgenfreii1
Abra (Abra) antwerpiensis1 Glibert 1945 abra
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Parvicardium straeleni1, Papillicardium papillosum1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Papillicardium papillosum1 Poli 1781 cockle
 Lucinida - Lucinidae
Anodontia (Loripinus) noerregaardi1, "Lucinoma borealis" = Lucinoma (Lucinoma) borealis1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata
 Pectinida - Pectinoidae
Korobkovia woodi1, Pseudamussium lilli1, Aequipecten angelonii1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Limidae
Limatula sulcata1
Limatula sulcata1 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Ostreida - Ostreidae
? Ostrea cf. edulis1, Cubitostrea digitalina1
? Ostrea cf. edulis1 Linnaeus 1758 oyster
Cubitostrea digitalina1 Eichwald 1830 oyster
 Arcida - Arcidae
Bathyarca pectunculoides1, Scapharca (Scapharca) diluvii1
Bathyarca pectunculoides1 Scacchi 1834 clam
Scapharca (Scapharca) diluvii1 Lamarck 1805 clam
 Arcida - Limopsidae
Limopsis (Limopsis) aurita1, "Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
 Terebratulida - Terebratulidae
Terebratula sp.1
Terebratula sp.1 Müller 1776
Bryozoa
  -
? Bryozoa indet.1
? Bryozoa indet.1 Ehrenberg 1831
Gymnolaemata
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Membraniporidae
Discoporella umbellata1, Reussirella haidingeri1
Discoporella umbellata1 Defrance 1823
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum, Flabellum tuberculatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
F. cristatum for. dietrichi
Flabellum tuberculatum Keferstein 1859 stony coral
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007