PBDB Collection

Santana do Cariri (general) (Cretaceous of Brazil)

Where: Ceará, Brazil (7.2° S, 39.7° W: paleocoordinates 8.6° S, 8.1° W)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Romualdo Formation (Santana Group), Late/Upper Aptian (122.5 - 113.0 Ma)

• Santana Formation suggested to be early Albian in age by Heimhofer & Hochuli (2010) on the basis of palynomorphs

•From Melo et al. 2020: In summary, this integrated study of foraminifera, ostracodes and other microfossil data from the Romualdo Formation indicates that the local Alagoas Stage (Ostracoda Zone RT-011) can now be constrained to the Aptian. Arai and Assine 2020 also support that the Romualdo Formation and Santana Group is entirely Aptian in age.

• member-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; concretionary, calcareous shale

• "lagoonal" (Kellner and Campos 2002); "The unit was deposited under quasi-marine conditions in the Araripe Basin, a large fault-bounded pull-apart basin that developed during the early stages of the opening of the Atlantic Ocean".

• fossils are from "limestone concretions embedded in shales" (Kellner and Campos 2002). "The Romualdo Member is a 4-30 m thick sequence of fissile, laminated black shales with numerous elliptical and sub-spherical concretions"

Size class: macrofossils

Preservation: concretion

Collection methods: quarrying, chemical,

• pterosaur is at the Museu de Ciencias da Terra/Departamento Nacional de Producao Mineral (DGM)

•SMNK: Staatliches Museum fuer Naturkunde, Karlsruhe, Germany

Primary reference: R. C. Martins Neto and A. W. A. Kellner. 1988. Primeiro registro de pena na Formacao Santana (Cretaceo inferior), Bacia do Araripe, nordeste do Brasil. Anais da Academia Brasileira de Ciencias 60:61-68 [J. Hunter/J. Hunter/J. Hunter]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 67712: authorized by John Alroy, entered by John Alroy on 06.12.2006, edited by Mark Uhen, Matthew Carrano, Richard Butler, Roger Benson and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Gnathostomata
-	Osteichthyes indet.⁵ Osteichthyes indet.⁵ bony fish "fish scales"
Aves
-	Aves indet. Aves indet. Linnaeus 1758 bird
Reptilia
Theropoda -	Santanaraptor placidus n. gen. n. sp.⁵ Santanaraptor placidus n. gen. n. sp.⁵ Kellner 1999 coelurosaur MN 4802-V
Theropoda - Spinosauridae	Spinosauridae indet.¹ Spinosauridae indet.¹ Stromer 1915 tetanuran theropod
Pterosauria - Thalassodromidae	Thalassodromeus sethi n. gen. n. sp.⁷, Tupuxuara sp.¹¹, Tupuxuara cf. longicristatus¹⁰, Tupuxuara leonardii n. sp.⁶, Tupuxuara deliradamus n. sp.¹⁴ Thalassodromeus sethi n. gen. n. sp.⁷ Kellner and Campos 2002 pterosaur DGM 1476-R, skull Tupuxuara sp.¹¹ Kellner and Campos 1988 pterosaur SMNK PAL 4330, partial skull and two cervical vertebrae Tupuxuara cf. longicristatus¹⁰ Kellner and Campos 1988 pterosaur SMNK PAL 3858, skull fragment Tupuxuara leonardii n. sp.⁶ Kellner and Campos 1994 pterosaur MCT 1495-R, anterior portion of skull Tupuxuara deliradamus n. sp.¹⁴ Witton 2009 pterosaur SMNK PAL 6410 (holotype), KPMNH DL 84, partial skulls
Pterosauria -	Anhanguera piscator n. sp.⁸, "Coloborhynchus robustus" = Anhanguera robustus², Unwindia trigonus n. gen. n. sp.⁹ Anhanguera piscator n. sp.⁸ Kellner and Tomida 2000 pterosaur NSM-PV 19892, partial skeleton "Coloborhynchus robustus" = Anhanguera robustus² Wellnhofer 1987 pterosaur SMNK 2302 PAL, anterior parts of upper and lower jaws Unwindia trigonus n. gen. n. sp.⁹ Martill 2011 pterosaur SMNK PAL 6597, partial rostrum with teeth
Testudines - Bothremydidae	Cearachelys placidoi n. gen. n. sp.³ Cearachelys placidoi n. gen. n. sp.³ Gaffney et al. 2001 sideneck turtle Holotype: MPSC a partial skull, shell, cervicals, and limb elements. Referred specimen: TUTg 1798—Nearly complete skeleton with skull, lower jaw, shell, cervicals, and limb elements (form 'near Santana do Cariri')
Testudines -	Caririemys violetae n. gen. n. sp.¹³ Caririemys violetae n. gen. n. sp.¹³ Oliveira and Kellner 2007 sideneck turtle partial skeleton consisting of an incomplete carapace, part of the last cervical vertebra, dorsal and caudal vertebrae, right pelvis and femur. The specimen is housed at the Museu Nacional (MN) of the Federal University of Rio de Janeiro (UFRJ) under the number MN 6919–V.
Testudines - Araripemydidae	Araripemys barretoi¹² Araripemys barretoi¹² Price 1973 sideneck turtle The fossils described below were obtained as nodules from Brazil via Herbert Axelrod: AMNH 22550; AMNH 24452; AMNH 24453; AMNH 24454; AMNH 22551; AMNH 24455; AMNH 24456; AMNH 24457, AMNH 24458, AMNH 24459, AMNH 24460, and AMNH 24461; Gaffney et al. 2006 additionally referred THUg 1907, skull with cervicals and shell fragments (purchased from Schwickert (Kranz)
Testudines - Protostegidae	Santanachelys gaffneyi n. gen. n. sp.⁴ Santanachelys gaffneyi n. gen. n. sp.⁴ Hirayama 1998 sea turtle
Deuteropoda
-	Ostracoda indet.⁵ Ostracoda indet.⁵ Latreille 1802 ostracod

¹Bittencourt and Kellner 2004; ²Fastnacht 2001; ³Gaffney et al. 2001; ⁴Hirayama 1998; ⁵Kellner 1999; ⁶Kellner and Campos 1994; ⁷Kellner and Campos 2002; ⁸Kellner and Tomida 2000; ⁹Martill 2011; ¹⁰Martill and Naish 2006; ¹¹Martill and Witton 2008; ¹²Meylan 1996; ¹³Oliveira and Kellner 2007; ¹⁴Witton 2009