Haikou [Yuanshan Fm (formerly Qiongzhusi)] (Cambrian of China)

Also known as Chengjiang biota; Jingmacun, Kunming

Where: Yunnan, China (24.8° N, 102.6° E: paleocoordinates 35.1° N, 161.1° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Yuanshan Formation, Atdabanian (520.0 - 516.0 Ma)

• LITHOSTRATIGRAPHY: Formerly from the Yuanshan Mbr, Qiongzhusi Fm. Yuanshan is raised to formation rank in recent publications (Steiner 2005, 2007). Wade-Giles romanization of Qiongzhusi is Chiungchussu. AGE: Eoredlichia Zone. Refined to Atdabanian by A. Hendy (2/2009)

• group of beds-level stratigraphic resolution

Environment/lithology: marine; lithified, gray, green, yellow, carbonaceous mudstone

• "benthic sediment flows... are believed to have been the the principal method of burial" (Shu et al. 1999b)

•"presence of agglutinated quartz grains, substantially coarser than the surrounding matrix... suggests... that the animal inhabited a higher-energy, sandy sea floor and was transported" (Shu et al. 2001)

• LITHOLOGY: Dark grey-green (yellow, when weathered), finely laminated mudstones, containing organic hash in great quantity. LITHIFICATION: Lithified, on the basis of figures.

Size class: macrofossils

Preservation: adpression, soft parts

Collection methods: surface (in situ), hydrochloric,

• COLLECTOR: Multiple collectors. REPOSITORY: Not stated in text.

Primary reference: M. Steiner, M. Zhu, Y. Zhao and B.-D. Erdtmann. 2005. Lower Cambrian Burgess Shale-type fossil associations of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 220:129-152 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 68083: authorized by John Alroy, entered by John Alroy on 02.01.2007, edited by Austin Hendy and Marcus Domansky

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for all groups. NOMENCLATURE: From a combination of authoritative and review publications, with modern nomenclature and species-resolution identifications.
unclassified
  -
Paraselkirkia
  -
Corynetis brevis
  -
Corynetis brevis Luo and Hu 1999
Sicyophorus
  -
Sicyophorus rara Luo and Hu 1999
Acosmia maotiania
  -
Acosmia maotiania Chen and Zhou 1997
 Hallucishaniids - Luolishaniidae
Luolishania longicruris Hou and Chen 1989
 Hallucishaniids -
Hallucigenia fortis1 Hou and Bergström 1995
Cardiodictyon
  -
Cardiodictyon catenulum Hou et al. 1991
Diania cactiformis
  -
Diania cactiformis n. gen. n. sp.2
Diania cactiformis n. gen. n. sp.2 Liu et al. 2011
Jianfengia
  -
 Parvancorinomorpha - Parvancorinidae
Primicaris larvaformis Zhang et al. 2003
Odaraia
  -
Megacheira
 Fortiforcipida - Fortiforcipidae
Fortiforceps foliosa Hou and Bergström 1997
Ercaia
  -
 Radiodonta - Amplectobeluidae
 Radiodonta - Anomalocarididae
Parapeytoia yunnanensis Hou et al. 1995
Anomalocaris saron Hou et al. 1995
Leanchoilia
  -
Megadictyon
  -
 Bradoriida -
Cindarella eucalla
  -
Cindarella eucalla Chen et al. 1996
Trilobita
 Redlichiida - Redlichiidae
Eoredlichia sp.5 Chang 1950 trilobite
Eoredlichia intermedia Lu 1940 trilobite
Kuanyangia pustulosa Lu 1941 trilobite
 Ptychopariida - Yunnanocephalidae
Yunnanocephalus sp.5 Kobayashi 1936 trilobite
Yunnanocephalus yunnanensis Mansuy 1912 trilobite
Crustacea
 Waptiida - Waptiidae
Waptia sp.5 Walcott 1912
 Protocaridida -
Sinoburius lunaris
  -
Sinoburius lunaris Hou et al. 1991
Artiopoda
 Conciliterga -
Kuamaia lata Hou 1987 artiopod
Saperion glumaceum Hou et al. 1991 artiopod
  -
Squamacula clypeata Hou and Bergström 1997 artiopod
Retifacies abnormalis
  -
Retifacies abnormalis Hou et al. 1989
 Fuxianhuiida - Fuxianhuiidae
Isoxys
  -
Isoxys auritus Jiang 1982
Isoxys curvirostratus Vannier and Chen 2000
Isoxys paradoxus Hou 1987
Urokodia
  -
Urokodia aequalis Hou et al. 1989
Xandarella
  -
Xandarella sp. Hou et al. 1991
 Nektaspida -
Naraoia sp.5 Walcott 1912
Naraoia longicaudata Zhang and Hou 1985
Naraoia spinosa Zhang and Hou 1985
Tanglangia
  -
 Bilateria -
Palaeoscolecida
 Cricocosmida - Tylotitidae
Tylotites petiolaris Luo and Hu 1999
 Cricocosmida - Cricocosmiidae
 Cricocosmida - Palaeoscolecidae
"Palaeoscolex sinensis" = Mafangscolex sinensis
"Palaeoscolex sinensis" = Mafangscolex sinensis Hou and Sun 1988
 Cricocosmida - Maotianshaniidae
? Maotianshania cylindrica Sun and Huo 1987
Hyolitha
 Hyolithida - Sulcavitidae
Lingulata
 Lingulida - Disciniidae
Heliomedusa orienta6 Sun and Hou 1987
Heliomedusa orienta Sun and Hou 1987
 Lingulida - Obolidae
"Lingulella chengjiangensis" = Eoglossa chengjiangensis
"Lingulella chengjiangensis" = Eoglossa chengjiangensis Jin et al. 1993
 Lingulida - Lingulellotretidae
 Lingulida - Botsfordiidae
Diandongia pista Rong 1974
 Lophophorata -
Cheungkongella ancestralis n. gen. n. sp.3 Shu et al. 2001
Cheungkongella ancestralis Shu et al. 2001
Phlogites longus Luo and Hu 1999
Cotyledion tylodes
  -
Cotyledion tylodes Luo and Hu 1999
Agnatha
  -
Haikouichthys ercaicunensis n. gen. n. sp.5 Luo et al. 1999 jawless fish
Haikouichthys ercaicunensis Luo et al. 1999 jawless fish
Myllokunmingia fengjiaoa n. gen. n. sp.5 Wagner 2017 jawless fish
Myllokunmingia fengjiaoa Wagner 2017 jawless fish
Xianguangia
  -
Xianguangia sinica Chen and Erdtmann 1991
Banffia
  -
Amiskwia
  -
Facivermis yunnanicus
  -
Facivermis yunnanicus Hou and Chen 1989
Facivermis
  -
"Xishania longiusula" = Facivermis longiusula
"Xishania longiusula" = Facivermis longiusula Hu 2002
Nectocaris
  - Nectocarididae
Petalilium latus Luo and Hu 1999
Demospongea
 Protomonaxonida - Leptomitidae
Paraleptomitella dictyodroma Chen et al. 1989 demosponge
Leptomitus teretiusculus Chen et al. 1989 demosponge
 Protomonaxonida - Hazeliidae
Hazelia palmata Walcott 1920 demosponge
 Protomonaxonida - Halichondritidae
Halichondrites sp. Walcott 1920 demosponge
 Protomonaxonida - Hamptoniidae
Hamptonia bowerbanki Walcott 1920 demosponge
 Protomonaxonida - Choiidae
Allantospongia mica Rigby and Hou 1995 demosponge
Choiaella radiata Rigby and Hou 1995 demosponge
Choia sp. Walcott 1920 demosponge
Hexactinellida
 Reticulosa - Protospongiidae
Saetaspongia densa Mehl and Reitner 1993 glass sponge
Stellostomites eumorphus
  -
Stellostomites eumorphus Sun and Hou 1987
Xidazoon stephanus
  -
Xidazoon stephanus n. gen. n. sp.4 Shu et al. 1999
Xidazoon stephanus Shu et al. 1999
Cambrorhytium
  -
Vetulicolida
  - Vetulicolidae
 Vetulicolata - Didazoonidae
Conicula striata
  -
Conicula striata Luo et al. 1999
Sandaokania latinodosa
  -
Sandaokania latinodosa Luo and Hu 1999
Lagenula striolata
  -
Lagenula striolata Luo and Hu 1999
Discoides abnormis
  -
Discoides abnormis Luo et al. 1999
Sinocylindra yunnanensis
  -
Sinocylindra yunnanensis Chen and Erdtmann 1991