PBDB Collection

Mont Berru (Paleocene of France)

Also known as Berru bed 5; Mouras Quarry; Cernay

Where: Champagne-Ardenne, France (49.3° N, 4.2° E: paleocoordinates 43.8° N, 1.7° E)

• coordinate estimated from map

When: Sables de Bracheux Formation, Thanetian (59.2 - 56.0 Ma)

• younger than carrière Lemoine, which is late Thanetian (Russell 1981); NP9, MP6 (Smith and Smith 2003)

•

•bed 5 (Hooker and Russell 2012)

• bed-level stratigraphic resolution

Environment/lithology: fluvial; conglomerate

• "sédiments fluviatiles"

• conglomerate (Smith and Smith 2003)

Size class: macrofossils

Primary reference: D. E. Russell. 1981. Un primate nouveau du Paléocène supérieur de France. Geobios 14(3):399-405 [J. Alroy/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 68428: authorized by Mark Uhen, entered by Mark Uhen on 22.01.2007, edited by Philip Mannion, Lars van den Hoek Ostende, John Alroy and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Amphibia
Proteida - Batrachosauroididae	Palaeoproteus gallicus⁴ Palaeoproteus gallicus⁴ Estes et al. 1967 salamander MNHN CR 6705-6708, 6722 (atlantes), 6709-6711 (trunk vertebrae) and 6712 (dentary)
Aves
Neornithes - Remiornithidae	Remiornis heberti¹⁴ Remiornis heberti¹⁴ Lemoine 1881 bird Referred material (MNHN BR)
Gastornithiformes - Gastornithidae	Gastornis minor¹⁴, Gastornis parisiensis¹⁴, Gastornis russelli n. sp.¹⁴ Gastornis minor¹⁴ Lemoine 1878 bird Referred material Gastornis parisiensis¹⁴ Hébert 1855 bird Referred material (MNHN BR) Gastornis russelli n. sp.¹⁴ Martin 1992 bird MNHN R 3560 - type (left tarsometatarsus); MNHN R 2583 (a bill fragment) may also belong to this species (Martin 1992)
Strigiformes -	Berruornis orbisantiqui n. gen. n. sp.¹⁶ Berruornis orbisantiqui n. gen. n. sp.¹⁶ Mourer-Chauviré 1994 owl MNHN R4155 - holotype (incomplete right tarsometatarsus); referred material: MNHN BR 11186 (distal left tibiotarsus), BR 11195 (proximal right tarsometatarsus), BR 12482-12483 (right tarsometatarsus) and BR 14571 (right tarsometatarsus)
Reptilia
Crocodylia -	Asiatosuchus depressifrons³, Diplocynodon remensis n. sp.¹⁵ Asiatosuchus depressifrons³ Blainville 1855 crocodilian Diplocynodon remensis n. sp.¹⁵ Martin et al. 2014 crocodilian MHNH F BR 4020 - type (a nearly complete skull); referred material (MNHN)
Testudinata - Sichuanchelyidae	Laurasichersis relicta n. gen. n. sp.¹⁹ Laurasichersis relicta n. gen. n. sp.¹⁹ Pérez-García 2020 turtle MNHN.F BR13513 - holotype
Testudines - Trionychidae	Trionyx sp.² Trionyx sp.² Geoffroy Saint-Hilaire 1809 softshell turtle
Testudines - Macrobaenidae	Gallica lapparentiana n. gen. n. sp.²⁰ Gallica lapparentiana n. gen. n. sp.²⁰ Pérez-García 2020 turtle MNHN.F BR13516 - holotype
Testudinata - Compsemydidae	Berruchelus russelli n. gen. n. sp.¹⁸ Berruchelus russelli n. gen. n. sp.¹⁸ Pérez-García 2012 turtle MNHN.F.BR 9110 - holotype (nuchal plate); referred material (MNHN): neurals (BR 2780, BR 9097, BR 9098, BR 15256), costals (BR 2827, BR 4181, BR 4288, BR 9038, BR 9043, BR 9062, BR 9092, BR 9101–BR 9103, BR 9105, BR 9112, BR 9115, BR 9118, BR 13430, BR 13433–BR 13647, BR 13651, BR 13652, BR 13713, BR 15002, BR 15005, BR 15009, BR 15043, BR 15059, BR 15151, BR 15157–BR 15159, BR 15169, BR 15408, BR 17468–BR 17470), peripherals (BR 872, BR 2676, BR 2690, BR 2775, BR 2776, BR 2793, BR 2834, BR 2861, BR 2728, BR 2734, BR 3655, BR 3658, BR 4182, BR 4217, BR 4218, BR 4289, BR 9093, BR 9099, BR 9100, BR 9104, BR 9109, BR 9113, BR 9114, BR 9116, BR 9117, BR 13447, BR 13455–BR 13457, BR 13709, BR 13728, BR 15010, BR 15126, BR 15255), suprapygal (BR 9096), pygal (BR 15010), epiplastra (BR 9091, BR 13454, BR 13725, BR 17471), entoplastra (BR 4206, BR 4212, BR 13726, BR 15010), hyoplastra (BR 1657, BR 4221, BR 9040, BR 9041, BR 9042, BR 9056, BR 9106, BR 13648, BR 13650, BR 15000, BR 15001, BR 15189, BR 40309), mesoplastra (BR 9036, BR 13649, BR 15003), hypoplastra (BR 2696, BR 2746, BR 4189, BR 4193, BR 4196, BR 9035, BR 9037, BR 9039, BR 9057, BR 9094, BR 9107), xiphiplastra (BR 13723, BR 15004), and seven fragments of indeterminate plates, numbered collectively as BR 15004; material originally preliminarily assigned to Compsemys (Broin 1977)
Mammalia
Macroscelidea - Louisinidae	Louisina mirabilis n. gen. n. sp.¹⁰, Louisina marci¹⁰, Thryptodon brailloni¹⁰, Berrulestes poirieri n. sp.¹⁰, Berrulestes phelizoni n. gen. n. sp.¹⁰, Berrulestes pellouini n. sp.¹⁰, Dipavali petri¹⁰, Walbeckodon girardi¹⁰, Gigarton sigogneauae¹⁰, Gigarton meyeri¹⁰ Louisina mirabilis n. gen. n. sp.¹⁰ Russell 1964 elephant shrew MNHN.F.BR143 - holotype (left dentary with crowns of M1–3 and alveoli of P2–4); paratypes: right dentary fragment with M2–3 (MNHN.F.BR-4406); referred material: right maxilla, M1–2, MNHN.F.BR-7-Lass; RM1, MNHN.F.BR-3-L; LM1, MNHN.F.I-729; RM3, MNHN. F.I-728 Louisina marci¹⁰ Hooker and Russell 2012 elephant shrew Two RM1, MNHN.F.BR-13-Pn, I-683; five LM2s, MNHN.F.I-583, I-709, I-714, I-718, BR-12-Bn; RM2, MNHN.F.I-707, LM3, IRSNB.CR-48-B-Wouters; RP3, MNHN.F.I-686; LP4, IRSNB.CR-34-B-Wouters; LM1, MNHN.F.I-721; two RM2s, MNHN.F.I-705, IRSNB. CR-42-B-Wouters Thryptodon brailloni¹⁰ Hooker and Russell 2012 elephant shrew Right M1, MNHN.F.I-681; RM2, MNHN.F.I-685; tentatively referred material: RP3, MNHN.F.I-691 Berrulestes poirieri n. sp.¹⁰ Hooker and Russell 2012 elephant shrew MNHN.F.I-725 - type (RM1); paratypes: RP4, MNHN.F.I-573; LM2, MNHN.F.BR-17478; two RM2s, MNHN.F.I-688, I-693; RM3, I-261; referred material: RM3, MNHN.F.I-713 Berrulestes phelizoni n. gen. n. sp.¹⁰ Hooker and Russell 2012 elephant shrew MNHN.F.BR-29-Ph - holotype (right dentary with P3–M3 and separate condyle); paratypes: RM1, MNHN.F.BR-6-L; LM2, MNHN.F.I-283; RP4, MNHN.F.I-602; RM1, MNHN.F.I-710; LM3, MNHN.F.BR-125-L; three RM3s, MNHN.F.I-580, I-754, I-247; referred material: RM1, MNHN.F.I-606; LM2, MNHN.F.I-692; RM2, MNHN.F.I-689 Berrulestes pellouini n. sp.¹⁰ Hooker and Russell 2012 elephant shrew MNHN.F.I-567 - type (LM1); paratypes: LM2, MNHN.F.I-253; two LM3s, MNHN.F.BR-3, BR-6 Dipavali petri¹⁰ Russell 1964 elephant shrew Associated RM1–3, MNHN.F. BR-15695–7; three RM1, MNHN.F.BR-2-L, BR-4-L, I-272; two LM2s, MNHN.F.I-587, I-690; LM3, MNHN.F.BR-14; two LP3s, MNHN.F.I-668, I-672; LP4, MNHN.F.I-671; twoRP4s, MNHN.F.BR-1-Ph, BR-9-Bn; LM1, MNHN. F.I-667; RM1, MNHN.F.I-673; L dentary, M2–3, MNHN.F.BR-1198; R dentary, M2–3, MNHN.F.BR-101; RM2, MNHN.F.BR-36-L; RM3, MNHN.F.I-669; RDP3, MNHN.F.BR-79 Walbeckodon girardi¹⁰ Hooker and Russell 2012 elephant shrew Right dentary with M1–2, MNHN.F.I-207 Gigarton sigogneauae¹⁰ Hooker and Russell 2012 elephant shrew LM2, MNHN.F.I-682 Gigarton meyeri¹⁰ Hooker and Russell 2012 elephant shrew LM1, MNHN.F.I-251
Condylarthra - Pleuraspidotheriidae	Orthaspidotherium edwardsi n. gen. n. sp.¹³ Orthaspidotherium edwardsi n. gen. n. sp.¹³ Lemoine 1885 placental
Condylarthra - Hyopsodontidae	Tricuspiodon sobrinus n. sp.²¹, Tricuspiodon magistrae n. sp.²¹ Tricuspiodon sobrinus n. sp.²¹ Russell 1980 condylarth BR-2-L - type Tricuspiodon magistrae n. sp.²¹ Russell 1964 condylarth BR 4426 - type
Primates - Toliapinidae	Sarnacius gingerichi¹¹ Sarnacius gingerichi¹¹ Russell 1981 primate MNHN BR-3-Ph
Primates -	Berruvius lasseroni n. gen. n. sp. Berruvius lasseroni n. gen. n. sp. Russell 1964 primate MNHN BR-141 - type; referred material: MNHN BR-8, BR-1109 and BR-22-BN
Primates - Plesiadapidae	Platychoerops antiquus n. sp.¹, Plesiadapis berruensis n. sp.¹², Plesiadapis tricuspidens⁸, Plesiadapis remensis⁸, Chiromyoides campanicus⁷ Platychoerops antiquus n. sp.¹ Boyer et al. 2012 primate NMB Bru4 - type (right lower p3-m3) Plesiadapis berruensis n. sp.¹² Jehle et al. 2018 primate Plesiadapis tricuspidens⁸ Gervais 1877 primate numerous specimens from the MNHN, YPM, UCMP, and other collections including several skulls Plesiadapis remensis⁸ Lemoine 1887 primate single MNHN specimen from Berru Chiromyoides campanicus⁷ Stehlin 1916 primate
Placentalia -	"? Lipotyphla indet." = Eulipotyphla⁵ "? Lipotyphla indet." = Eulipotyphla⁵ placental lower molars
Acreodi - Mesonychidae	Dissacus europaeus²³ Dissacus europaeus²³ Lemoine 1891 condylarth IRSNB M2096, MNHN.F.2003–2.1121, MNHN.F.BR 13937, 1069, 12524, 1033
Procreodi - Arctocyonidae	Landenodon phelizoni n. sp.²¹, Landenodon lavocati²¹, Arctocyonides arenae²² Landenodon phelizoni n. sp.²¹ Russell 1980 condylarth BR 10156 - type Landenodon lavocati²¹ Russell 1980 condylarth BR-13-Bn Arctocyonides arenae²² Russell 1964 condylarth either Cernay or Berru
Perissodactyla -	"cf. Microhyus sp." = Teilhardimys²⁴, Teilhardimys brisswalteri¹⁰ "cf. Microhyus sp." = Teilhardimys²⁴ Kretzoi and Kretzoi 2000 odd-toed ungulate MNHN L−3−BR (right M1) Teilhardimys brisswalteri¹⁰ Hooker and Russell 2012 odd-toed ungulate Paratypes: RP4, MNHN.F.I-124; RM1, MNHN.F.I-712; two LM2s, MNHN.F.I-722, I-730; LP3, MNHN.F.I-708; RM1, MNHN.F.I-706; LM2, MNHN.F.I-242; two RM2s, MNHN.F.I-244, I-716; LM3, MNHN.F.I-715; two RM3s, MNHN.F.I-582, I-720; referred material: LM2, MNHN.F.I-726; LM1, BR-39
Theriamorpha - Adapisoriculidae	Bustylus cernaysi⁶, Afrodon sp.⁵ Bustylus cernaysi⁶ Gheerbrant and Russell 1991 eutherian MNHN BR 4 Bn, BRL 8 P, BR 9 Ph Afrodon sp.⁵ Gheerbrant 1988 eutherian e.g. MNHN BR-9-Ph
Theriamorpha - Pseudorhyncocyonidae	Phakodon levei⁹ Phakodon levei⁹ Russell et al. 1967 eutherian MNHN.F.I-601
Multituberculata - Kogaionidae	Hainina vianeyae n. sp.¹⁷ Hainina vianeyae n. sp.¹⁷ Peláez-Campomanes et al. 2000 multituberculate either Cernay or Berru

¹Boyer et al. 2012; ²Broin 1977; ³Delfino et al. 2019; ⁴Estes et al. 1967; ⁵Gheerbrant and Russell 1989; ⁶Gheerbrant and Russell 1991; ⁷Gingerich 1973; ⁸Gingerich 1976; ⁹Hooker 2013; ¹⁰Hooker and Russell 2012; ¹¹Hooker et al. 1999; ¹²Jehle et al. 2018; ¹³Ladevèze et al. 2010; ¹⁴Martin 1992; ¹⁵Martin et al. 2014; ¹⁶Mourer-Chauviré 1994; ¹⁷Peláez-Campomanes et al. 2000; ¹⁸Pérez-García 2012; ¹⁹Pérez-García 2020; ²⁰Pérez-García 2020; ²¹Russell 1980; ²²Smith and Smith 2003; ²³Solé et al. 2018; ²⁴Tabuce et al. 2006