PBDB Collection

Karyidia (Miocene of Greece)

Also known as Karydia I + II (KAR)

Where: Thrace, Greece (41.0° N, 25.3° E: paleocoordinates 41.4° N, 24.0° E)

• coordinate estimated from map

When: MN 4 (16.9 - 16.0 Ma)

Environment/lithology: terrestrial; lithology not reported

Size classes: macrofossils, mesofossils

Reposited in the AMPG

Primary reference: C. Doukas and L.W. van den Hoek Ostende. 2006. Insectivores (Erinaceomorpha, Soricomorpha; Mammalia)from Karydia and Komotini (Thrace, Greece; MN 4/5). Beiträge zur Paläontologie 30:109-131 [L. van den Hoek Ostende/L. van den Hoek Ostende]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 69435: authorized by Lars van den Hoek Ostende, entered by Lars van den Hoek Ostende on 26.02.2007, edited by Evangelos Vlachos and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Amphibia
Salientia -	Anura indet.² Anura indet.² Fischer von Waldheim 1813 frog KR2: a le humerus (UU KR2 5015) and a fragment of radioulna (UU KR2 5016). KR3: a premaxilla (AMPG KR3 003), an ilium (AMPG KR3 024), and eight phalanxes (AMPG KR3 006 and AMPG KR3 023).
Salientia - Discoglossidae	Latonia cf. gigantea² Latonia cf. gigantea² Lartet 1851 painted frog KR3: 15 maxillae (AMPG KR3 004, AMPG KR3 005, AMPG KR3 014–AMPG KR3 016, AMPG KR3 027, AMPG KR3 030, AMPG KR3 048–AMPG KR3 055), a frontoparietal (AMPG KR3 038), two praearticulars (AMPG KR3 028 and AMPG KR3 029), two trunk vertebrae (AMPG KR3 031 and AMPG KR3 033), a sacral vertebra (AMPG KR3 032), and four ilia (AMPG KR3 011, AMPG KR3 026, AMPG KR3 044, and AMPG KR3 045).
Caudata - Urodela	Urodela indet.² Urodela indet.² Duméril 1805 salamander KR3: a tibia (AMPG KR3 037).
Mammalia
Soricomorpha - Plesiosoricidae	Plesiosorex sp. Plesiosorex sp. Pomel 1848 placental
Soricomorpha - Talpidae	Desmanodon antiquus, Myxomygale cf. hutchisoni Desmanodon antiquus Ziegler 1985 mole Myxomygale cf. hutchisoni Ziegler 1985 mole
Soricomorpha - Soricidae	Heterosorex sp., Lartetium cf. dehmi, Paenelimnoecus sp. Heterosorex sp. Gaillard 1915 shrew Lartetium cf. dehmi shrew Paenelimnoecus sp. Baudelot 1972 red-toothed shrew
Tribosphenida - Erinaceoidae	Plesiodimylus aff. crassidens Plesiodimylus aff. crassidens Engesser 1980 placental
Tribosphenida - Erinaceidae	Galerix kostakii n. sp. Galerix kostakii n. sp. Doukas and van den Hoek Ostende 2006 hedgehog
Rodentia - Cricetidae	Eumyarion aff. latior³, Karydomys boskosi n. sp.⁴, Karydomys symeonidisi n. sp.³, Cricetodon aliveriensis³, cf. Deperetomys sp.³, "Mirabella tuberosa" = Mirrabella tuberosa³, Democricetodon franconicus³, Democricetodon cf. gaillardi³, "Democricetodon gracilis" = Democricetodon minor gracilis³ Eumyarion aff. latior³ Schaub and Zapfe 1953 rodent Karydomys boskosi n. sp.⁴ Theocharopoulos 2000 rodent Karydomys symeonidisi n. sp.³ Theocharopoulos 2000 rodent Cricetodon aliveriensis³ Klein Hofmeijer and de Bruijn 1988 rodent cf. Deperetomys sp.³ Mein and Freudenthal 1971 rodent "Mirabella tuberosa" = Mirrabella tuberosa³ de Bruijn et al. 1987 rodent Democricetodon franconicus³ Fahlbusch 1966 rodent Democricetodon cf. gaillardi³ Schaub 1925 rodent "Democricetodon gracilis" = Democricetodon minor gracilis³ Fahlbusch 1964 rodent
Rodentia - Muridae	Debruijnia kostakii n. sp.¹ Debruijnia kostakii n. sp.¹ de Bruijn 2017 blind mole
Rodentia - Gliridae	Miodyromys cf. praecox³, Glirulus (Glirulus) diremptus³, Glis galitopouli³, Seorsumuscardinus alpinus³, Glirudinus gracilis³, Glirulus (Paraglirulus) agelakisi³ Miodyromys cf. praecox³ Wu 1993 dormouse Glirulus (Glirulus) diremptus³ Mayr 1979 Japanese dormouse Glis galitopouli³ Van der Meulen and de Bruijn 1982 dormouse Seorsumuscardinus alpinus³ de Bruijn 1998 dormouse Glirudinus gracilis³ Dehm 1950 dormouse Glirulus (Paraglirulus) agelakisi³ Van der Meulen and De Bruijn 1982 dormouse
Rodentia - Anomalomyidae	Anomalomys aliveriensis³ Anomalomys aliveriensis³ Klein Hofmeijer and de Bruijn 1985 rodent
Rodentia - Eomyidae	Ligerimys sp.³ Ligerimys sp.³ Schaub 1951 rodent
Rodentia - Sciuridae	Blackia miocenica³, Aliveria luteijni³, Palaeosciurus aff. fissurae³ Blackia miocenica³ squirrel Aliveria luteijni³ squirrel Palaeosciurus aff. fissurae³ squirrel
Reptilia
Squamata -	Squamata indet.², Scincomorpha indet.² Squamata indet.² squamates KR3: a caudal vertebra (AMPG KR3 025) and an osteoderm (AMPG KR3 009). Scincomorpha indet.² squamates KR3: a dentary (AMPG KR3 010).
Squamata - Lacertidae	Lacertidae indet.² Lacertidae indet.² Bonaparte 1831 squamates KR3: two maxillae (AMPG KR3 017–AMPG KR3 018) and an isolated tooth (AMPG KR3 007).
Serpentes -	Serpentes indet.² Serpentes indet.² snake KR2: a cloacal or anterior caudal vertebra (UU KR2 5022), two fragmen- tary vertebrae (UU KR2 5023 and UU KR2 5024), and a cloacal or anterior caudal vertebra (UU KR2 5021). KR3: a fragment of a pterygoid (AMPG KR3 013), and two posterior caudal vertebrae (AMPG KR3 021 and AMPG KR3 022).
Squamata - Colubridae	Colubridae indet.² Colubridae indet.² Oppel 1811 colubrid snake KR2: a posterior trunk vertebra (UU KR2 5018), a posterior trunk vertebra (UU KR2 5019), a caudal vertebra (UU KR2 5025), and a fragmentary vertebra (UU KR2 5020). KR3: a frag- ment of a trunk vertebra (AMPG KR3 020).
Squamata - Anguidae	Anguinae indet.², cf. Ophisaurus sp.² Anguinae indet.² Gray 1825 squamates KR2: seven oste- oderms (UU KR2 5006–UU KR2 5013). KR3: 11 osteoderms (AMPG KR3 001, AMPG KR3 002, AMPG KR3 008, AMPG KR3 019, AMPG KR3 039–AMPG KR3 043, AMPG KR3 046, AMPG KR3 047, AMPG KR3 056, and AMPG KR3 057). cf. Ophisaurus sp.² Daudin 1803 glass lizard KR3: a tooth (AMPG KR3 035) and a trunk vertebra (AMPG KR3 034).
Testudines -	Testudines indet.² Testudines indet.² Batsch 1788 turtle KR2: three shell fragments (UU KR2 5001–UU KR2 5003).

¹de Bruijn 2017; ²Georgalis et al. 2017; ³Koufos 2006; ⁴Mörs and Kalthoff 2004