Where: Aisén, Chile (46.9° S, 72.3° W: paleocoordinates 47.7° S, 66.6° W)
• coordinate estimated from map
• local area-level geographic resolution
When: Santacrucian mammal zone, Galera Formation, Burdigalian (20.4 - 16.0 Ma)
• "a thick stratigraphic sequence, locally exposed as a prominent butte... approximately 300 m. thick sequence"; unit is "an unnamed formation, possibly equivalent to the Río Zeballos Formation"; mammals are from "Nine major mammal-bearing horizons... though the sequence is fossiliferous throughout"; much material is from "the rich quarry sites at the base of the terrestrial section (level E-O)"
•McGrath et al 2020: The mammal-bearing beds have been variably Buenos Aires) and Lago Cochrane (¼ Lago Pueyrred?on; assigned to the R?ıo Zeballos (R. H. Niemeyer 1975), Santa Cruz (de la Cruz et al. 2004), Galera (J. Niemeyer et al. 1984), or Pampa Castillo (Scalabrino 2009) formations (see discussion in Flynn et al. [2002] and Folguera et al. [2018]). Pending formal naming, designation of a type section, and a published descrip- tion of the Pampa Castillo formation, we continue to refer these beds to the Galera Formation, consistent with previous work (Flynn et al. 2002). The sequence has been correlated stratigraphically and temporally to the Santa Cruz Formation in Argentina (J. Niemeyer et al. 1984; Flynn et al. 2002). Folguera et al. (2018) recently provided a U/Pb date of 18.7 ± 0.3 Ma for zir- cons from a reworked tuff about 10 m above the contact between the terrestrial fossil-bearing unit and the grada- tionally and conformably underlying marine Guadal Formation at Meseta Guadal. A U/Pb age of ?18Ma from a tuff within the fossil-bearing unit (Su?arez et al. 2015) is consistent with the known age range of the Santacrucian SALMA and Santa Cruz Formation depos- its and faunas (Flynn & Swisher 1995; Croft et al. 2004; Cuiti~no et al. 2016; and discussion below). No litopterns have been reported from the Galera Formation (Bostelmann & Buldrini 2012). The Pampa Castillo at Pampa Guadal, ?20km north-west of Pampa Castillo fauna is referred to the Santacrucian SALMA, although only the rodents (Dodson 1994; Chick et al. 2010) and palaeothentids (Flynn et al. 2002) have been described comprehensively. This study represents the first detailed assessment of the fauna’s SANUs.
•McGrath et al. 2022: Geochronological age estimates suggest that Pampa Castillo is also closer temporally to “Pinturan” faunas than either is to core Santacrucian faunas. Folguera et al. (2018) reported a U/Pb date of 18.7 +/-0.3 Ma from a reworked tuff directly below strata bearing the Pampa Castillo fauna. “Pinturan” faunas of the lmPF and Upper Fossil Zone at Gran Barranca are ~19.0–17.9 Ma (Perkins et al., 2012; Dunn et al., 2013), whereas core Santacrucian faunas are ~17.5–15.6 Ma (Cuitiño et al., 2016; Trayler et al., 2020).
• formation-level stratigraphic resolution
Environment/lithology: fluvial; sandstone and siltstone
Size class: macrofossils
Collection methods: surface (float),
Primary reference: J. J. Flynn, M. J. Novacek, H. E. Dodson, D. Frassinetti, M. C. McKenna, M. A. Norell, K. E. Sears, C. C. Swisher, III, and A. R. Wyss. 2002. A new fossil mammal assemblage from the southern Chilean Andes: implications for geology, geochronology, and tectonics. Journal of South American Earth Sciences 15(3):285-302 [J. Alroy/J. Alroy]more details
Purpose of describing collection: biostratigraphic analysis
PaleoDB collection 71843: authorized by John Alroy, entered by John Alroy on 14.05.2007, edited by Miranta Kouvari, Grace Varnham and Kateryn Pino
Creative Commons license: CC BY (attribution)
Taxonomic list
Mammalia | |
Astrapotherium sp. Burmeister 1879 placental | |
Hegetotherium sp. Ameghino 1887 notoungulate | |
Protypotherium sp. Ameghino 1882 notoungulate | |
Homalodotherium sp. Huxley 1870 notoungulate | |
Adinotherium sp. Ameghino 1887 notoungulate
Nesodon sp. Owen 1846 notoungulate | |
Hapalops sp. Ameghino 1887 edentate | |
Proeutatus sp., "Prozaedius sp." = Prozaedyus
Proeutatus sp. Ameghino 1891 edentate
"Prozaedius sp." = Prozaedyus Ameghino 1891 edentate | |
"Propalaeohoplophorus sp." = Propalaehoplophorus
"Propalaeohoplophorus sp." = Propalaehoplophorus Ameghino 1887 edentate | |
Eosteiromys cf. annectens1, Steiromys sp., Steiromys sp.1, Steiromys duplicatus, Spaniomys riparius, Acarechimys constans3, Acarechimys minutus3, Acarechimys cf. minutissimus3, Caviocricetus guenekko n. sp.3
Eosteiromys cf. annectens1 Ameghino 1901 caviomorph Fossil Horizon E-0 (SGOPV 2157, 2688) and unknown level (SGOPV 2752)
Steiromys sp.1 Ameghino 1887 caviomorph Fossil Horizon E-0 (SGOPV 2689) and Fossil Horizon 7 (SGOPV 2190)
Steiromys duplicatus Ameghino 1887 caviomorph
Spaniomys riparius Ameghino 1887 caviomorph
Acarechimys constans3 Ameghino 1887 caviomorph Fossil Horizon E-0 W (SGOPV 2198, 2199, 2707), Fossil Horizon 7 (SGOPV 2135, 2137, 2138, 2139, 2143, 2705) and Bangles Quarry (SGOPV 2362)
Caviocricetus guenekko n. sp.3 McGrath et al. 2023 caviomorph Fossil Horizon E-0 W. Holotype: SGOPV 2194, maxillary fragment bearing right DP4-M1
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Sciamys principalis Ameghino 1887 caviomorph | |
Dudumus berggreni n. sp.3 McGrath et al. 2023 caviomorph Fossil Horizon 7. Holotype: SGOPV 2140, maxillary fragment bearing left M1–2
Stichomys ? sp. Ameghino 1887 caviomorph | |
Prostichomys "n. sp."3 Kramarz 2001 spiny rat sp. nov. (unnamed). Fossil Horizon E-0 E. SGOPV 2706, left m1 or 2
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Prolagostomus pusilus, Prolagostomus notatus, "Prolagostomus pusillus" = Prolagostomus pusillus1, "Prolagostomus profluens" = Prolagostomus pusillus, "Prolagostomus divisus" = Prolagostomus pusillus
Prolagostomus pusilus chinchilid
Prolagostomus notatus chinchilid
"Prolagostomus pusillus" = Prolagostomus pusillus1 Ameghino 1887 chinchilid Fossil Horizon E-2 (SGOPV 2612), Fossil Horizon 4 (SGOPV 2613), Fossil Horizon 6-7 (SGOPV 2427, 2607, 2608, 2609, 2610, 2611), and Fossil Horizon 7 (SGOPV 2599, 2600, 2601, 2602, 2603, 2604, 2605, 2606, 2614, 2615)
"Prolagostomus profluens" = Prolagostomus pusillus Ameghino 1887 chinchilid
"Prolagostomus divisus" = Prolagostomus pusillus Ameghino 1887 chinchilid | |
Luantus sp.1, Luantus minor1, Neoreomys australis, cf. Eocardia sp.1, "Eocardia perforata" = Eocardia robusta, cf. Eocardia excavata1
Luantus minor1 Pérez et al. 2010 caviomorph Fossil Horizon 4 (SGOPV 2691, 2697) and Fossil Horizon 7 (SGOPV 2134); Galera Formation, Pampa Castillo, Chile.
cf. Eocardia sp.1 Ameghino 1887 caviomorph Fossil Horizon 7: SGOPV 2695, partial left m2 and left m3 in mandibular fragment
"Eocardia perforata" = Eocardia robusta Vucetich 1984 caviomorph
cf. Eocardia excavata1 Ameghino 1891 caviomorph Fossil Horizon 7: SGOPV 2133, left M1–2; SGOPV 2690, left m1–2; SGOPV 2692, right M3; SGOPV 2693, left M1 or 2; SGOPV 2694, right M2; and SGOPV 2696, right M1
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Schistomys erro Ameghino 1887 caviomorph | |
Perimys "n. sp."1, Perimys intermedius1, "Perimys erutus" = Perimys erutus1, "Perimys procerus" = Perimys erutus, "Perimys scalaris" = Perimys erutus, "Perimys onustus" = Perimys onustus, "Perimys perpinguis" = Perimys onustus, "Perimys impactus" = Perimys onustus
Perimys "n. sp."1 Ameghino 1887 caviomorph Fossil Horizon E-0 (SGOPV 2285, 2401, 2650), Fossil Horizon E-2 (SGOPV 2267, 2686), Fossil Horizon 4 (SGOPV 2646, 2647, 2648, 2649, 2684), and Fossil Horizon 7 (SGOPV 2431, 2679)
Perimys intermedius1 Kramarz 2002 caviomorph Fossil Horizon E-0 (SGOPV 2299), Fossil Horizon 4 (SGOPV 2651, 2652, 2653), Fossil Horizon 4abc (SGOPV 2281), and Fossil Horizon 7 (SGOPV 2645, 2654)
"Perimys erutus" = Perimys erutus1 Ameghino 1887 caviomorph Fossil Horizon E-0 (SGOPV 2386, 2618, 2627, 2628, 2631, 2632, 2633, 2634, 2635, 2636, 2637, 2638, 2639, 2640, 2672, 2722), Fossil Horizon 1A (SGOPV 2361), Fossil Horizon E-2 (SGOPV 2616), Fossil Horizon 4 (SGOPV 2617, 2617, 2621, 2623, 2624, 2625, 2629, 2630), Fossil Horizon 7 (SGOPV 2641, 2642, 2643, 2644), “Glyptodont Glen” (SGOPV 2294, 2620, 2626), and unknown Fossil Horizon (SGOPV 2391, 2622)
"Perimys procerus" = Perimys erutus Ameghino 1887 caviomorph
"Perimys scalaris" = Perimys erutus Ameghino 1887 caviomorph
"Perimys onustus" = Perimys onustus Ameghino 1887 caviomorph Fossil Horizon E-0 (SGOPV 2421, 2673) Fossil Horizon E-2 (SGOPV 2101, 2675), Fossil Horizon 2 (SGOPV 2014), Fossil Horizon 3 (SGOPV 2030), Fossil Horizon 4 (SGOPV 2045, 2056, 2060, 2273, 2661, 2662, 2663, 2664, 2671, 2674, 2681, 2682), Fossil Horizon 5 (SGOPV 2080, 2665, 2687), Fossil Horizon 6-7 (SGOPV 2655, 2656, 2657, 2658, 2659, 2660), Fossil Horizon 7 (SGOPV 2121, 2378, 2666, 2667, 2668, 2669, 2670, 2677, 2678, 2680), “Glyptodont Glen” (SGOPV 2676), and unknown Fossil Horizon (SGOPV 2683)
"Perimys perpinguis" = Perimys onustus Ameghino 1887 caviomorph
"Perimys impactus" = Perimys onustus Ameghino 1887 caviomorph | |
Scleromys sp. Ameghino 1887 caviomorph
Scleromys quadrangulatus1 Kramarz 2006 caviomorph Fossil Horizon 4 (SGOPV 2699, 2700, 2702, 2703) and unknown Fossil Horizon (SGOPV 2701),
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Proterotheriidae indet.2, Picturotherium sp.2, Proterotherium sp., cf. Thoatherium sp.2, Thoatherium cf. minusculum2
Proterotherium sp. Ameghino 1883 placental | |
Theosodon cf. gracilis Ameghino 1891 placental | |
Palaeothentes smeti n. sp., Palaeothentes pascuali, Palaeothentes minutus, Palaeothentes intermedius
Palaeothentes smeti n. sp. Flynn et al. 2002 marsupial
Palaeothentes pascuali Bown and Fleagle 1993 marsupial
Palaeothentes minutus Ameghino 1887 marsupial
Palaeothentes intermedius Ameghino 1887 marsupial | |
Cladosictis sp. Ameghino 1887 metatherian |