PPP# 177 - Escudo de Veraguas [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 17625

Where: Bocas, Panama (9.1° N, 81.6° W: paleocoordinates 9.0° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999).

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 8/7/87. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 79771: authorized by Arnold Miller, entered by Kate Bulinski on 23.03.2008, edited by Austin Hendy

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Anthozoa
 Scleractinia - Caryophylliidae
Asterosmilia profunda Duncan 1864 stony coral
Gymnolaemata
 Cheilostomata - Mamilloporidae
 Cheilostomata - Membraniporidae
 Cheilostomata - Lepraliellidae
 Cheilostomata - Celleporidae
Turbicellepora sp. Ryland 1963
Scaphopoda
 Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758 tusk shell
sensu lato
 Gadilida - Gadilidae
Cadulus sp. Philippi 1844 tusk shell
Bivalvia
 Arcida - Glycymerididae
Tucetona sp. Iredale 1939 clam
 Arcida - Arcidae
Anadara (Rasia) sp. Gray 1857 ark
 Pectinida - Pectinoidae
Argopecten sp. Monterosato 1889 calico scallop
 Lucinida - Lucinidae
"Lucinisca sp." = Lucina (Lucinisca)
"Lucinisca sp." = Lucina (Lucinisca) Dall 1901 clam
 Nuculanida - Nuculanidae
Saccella sp. Woodring 1925 pointed nut clam
 Nuculida - Nuculidae
Leionucula sp. Quenstedt 1930 nut clam
Gastropoda
 Thecosomata - Limacinidae
Limacina sp. Bosc 1817 snail
 Heterostropha - Architectonicidae
Architectonica sp. Röding 1798 sundial
 Sorbeoconcha - Naticidae
"Natica (Naticarius) sp." = Naticarius
"Natica (Naticarius) sp." = Naticarius Dumeril 1806 moon snail
 Sorbeoconcha - Vermetidae
Serpulorbis sp. Sasso 1827 worm shell
 Neogastropoda - Mitridae
Subcancilla sp. Olsson and Harbison 1953 mitre shell
 Neogastropoda - Buccinidae
Metula sp. Adams and Adams 1853 true whelk
Antillophos (Antillophos) sp. Woodring 1928 true whelk
 Neogastropoda - Olividae
Olivella sp. Swainson 1831 olive snail
sensu lato
Olivella (Minioliva) sp. Olsson 1956 olive snail
Olivella (Macgintiella) sp. Olsson 1956 olive snail
Oliva (Oliva) sp. Bruguière 1789 olive snail
 Neogastropoda - Turridae
Scobinella sp. Conrad 1848 turrid
Polystira sp. Woodring 1928 turrid
 Neogastropoda - Conidae
Conus sp. Linnaeus 1758 cone shell
 Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891 auger snail
 Neogastropoda - Marginellidae
Volvarina sp. Hinds 1844 margin shell
 Sorbeoconcha - Atlantidae
Atlanta sp. Lesueur 1817 snail
 Pseudomelanoidea - Pseudomelaniidae
Crassispira (Crassispira) sp. Swainson 1840 snail