PBDB Collection

Vyazniki Localities, PIN 1100 (Permian of Russian Federation)

Also known as Klyazma River; Bykovka gully; Zelenaya Gorka; Bykovka lake; Bykovka; Vyazniki-1; Vyazniki-2; Metallist; Yartsevo; Tolmachevo; Balymotikha-1; Balymotikha-2

Where: Vladimir, Russian Federation (56.3° N, 42.1° E: paleocoordinates 28.8° N, 37.7° E)

• coordinate estimated from map

• local area-level geographic resolution

When: Archosaurus rossicus other zone, Vyatkian (259.5 - 257.4 Ma)

• Vyazniki Assemblage; the terminal part of the Late Tatarian (terminal

•part of the Vyatkian) (Golubev 2000; Benton 2012, Geology).

• group of beds-level stratigraphic resolution

Environment/lithology: "floodplain"; sandstone and claystone

• "These deposits were accumulated in a number of channels and abandoned channels (or oxbow-lakes) of the vast flood-plain of a great river system directed from east to west, that is, from the Ural Mountains to the plain." (Sennikov & Golubev, 2006)

• "All fossils were found in sands and clays of alluvial genesis" (Sennikov & Golubev, 2006)

Size classes: macrofossils, mesofossils

Reposited in the PIN

Primary reference: A. G. Sennikov and V. K. Golubev. 2006. Vyazniki Biotic Assemblage of the Terminal Permian. Paleontological Journal 40(suppl. 4):S475-S481 [J. Mueller/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 86346: authorized by Johannes Mueller, entered by Torsten Liebrecht on 31.01.2009, edited by Richard Butler and Bethany Allen

Creative Commons license: CC BY (attribution)

Taxonomic list

• The two elements listed as Obirkovia sp. by Bulanov & Yashina (2005) are identified as Elginia(?) sp. and Elginia sp. by Ivakhnenko (1997, in Ivakhnenko et al. [eds.] Permian and Triassic Tetrapods of Eastern Europe) and Ivakhnenko (2008), respectively. Since Ivakhnenko (2008) does not mention the re-assignment of these specimens to Obirkovia sp. by Bulanov & Yashina (2005), Ivakhnenko's (2008) assigment to Elginia sp. is ignored here.

Show authors, comments, and common names

unclassified
-	Clinocypris cf. elongata, Clinocypris sp., Darwinuloides ex gr. svijazhicus, Nerechtina cf. plana, Suchonella cf. posttypica, Suchonella ex gr. typica, Wjatkellina ? pseudoobliqua, Wjatkellina fragilina, Wjatkellina vladimirinae, Wjatkellina cf. vladimirinae Clinocypris cf. elongata Clinocypris sp. Darwinuloides ex gr. svijazhicus Nerechtina cf. plana Suchonella cf. posttypica Suchonella ex gr. typica Wjatkellina ? pseudoobliqua Wjatkellina fragilina Wjatkellina vladimirinae Wjatkellina cf. vladimirinae
Chondrichthyes
Ctenacanthiformes -	Wodnika sp. Wodnika sp. Münster 1843 elasmobranch
Amphibia
Temnospondyli - Dvinosauridae	Dvinosaurus egregius n. sp. Dvinosaurus egregius n. sp. Shishkin 1968 tetrapod PIN 1100/23 (holotype), skull, plus additional specimens
Osteichthyes
Seymouriamorpha - Kotlassidae	Kotlassia sp.⁷, Microphon aff. exiguous⁷, Microphon terminalis n. sp.³ Kotlassia sp.⁷ Amalitzky 1921 tetrapod Microphon aff. exiguous⁷ tetrapod Microphon terminalis n. sp.³ Bulanov 2020 tetrapod PIN 5078/18 (holotype), "a crown of an isolated maxillary tooth"; PIN 5078/11, "lacrimal"; PIN 5078/14, "squamosal"; PIN 5078/12, "postparietal"; PIN 5078/13, "tabular"; PIN 5078/6, "intertemporal"; PIN 5078/16, "parabasisphenoid"; 147 teeth
Seymouriamorpha - Karpinskiosauridae	Karpinskiosaurus sp.² Karpinskiosaurus sp.² Sushkin 1926 tetrapod PIN 1100/145, jugal
Reptilia
Procolophonomorpha - Elginiidae	Elginiidae indet.⁴, Obirkovia sp.⁴ Elginiidae indet.⁴ Cope 1896 parareptile PIN 1100/140, 1100/500 (two osteoderms of the skull) Obirkovia sp.⁴ Bulanov and Yashina 2005 parareptile PIN 1100/141 (incomplete left nasal), 142 (single osteoderm/dermal scute); both are from Vyazniki-2; see comments on taxonomic list for previous and subsequent identifications
Diapsida - Proterosuchidae	Archosaurus sp.⁵, Archosaurus rossicus n. gen. n. sp.⁵ Archosaurus sp.⁵ Tatarinov 1960 archosauromorph PIN 1100/78, 1100/427 Archosaurus rossicus n. gen. n. sp.⁵ Tatarinov 1960 archosauromorph PIN 1100/55 (type), 1100/66, 1100/67, 1100/67a, 1100/84, 1100/84a, 1100/85a
Osteichthyes
Dicynodontia - Dicynodontidae	Dicynodontidae indet. Dicynodontidae indet. Owen 1859 dicynodont
Therocephalia - Whaitsiidae	Megawhaitsia patrichae n. gen. n. sp.⁶, Moschowhaitsia vjuschkovi n. gen. n. sp.¹ Megawhaitsia patrichae n. gen. n. sp.⁶ Ivakhnenko 2008 therapsid PIN 1100/101 (type) Moschowhaitsia vjuschkovi n. gen. n. sp.¹ Tatarinov 1963 therapsid PIN 1100/20 (type; from Vyazniki-1); PIN 1100/19, 21, 30, 68, 74
Therocephalia - Nanictidopidae	Purlovia maxima⁷ Purlovia maxima⁷ Ivakhnenko 2011 therapsid PIN 1100/104
Tetrapoda - Bystrowianidae	Bystrowiana permira n. gen. n. sp.⁸ Bystrowiana permira n. gen. n. sp.⁸ Vjushkov 1957 tetrapod PIN 1100/1 (single vertebra with osteoderm plate)
Tetrapoda - Chroniosuchidae	Uralerpeton tverdochlebovae n. gen. n. sp.⁸ Uralerpeton tverdochlebovae n. gen. n. sp.⁸ Golubev 1998 tetrapod PIN 1100/8 (single osteoderm plate)
Actinopteri
Scanilepiformes - Evenkiidae	Toyemia sp., Toyemia blumentalis Toyemia sp. Minikh 1990 Toyemia blumentalis Minikh 1995
Osteichthyes
- Discordichthyidae	Mutovinia sp., Mutovinia stella, ? Geryonichthys sp. Mutovinia sp. Minikh 1992 "Mutovinia sp. nov." Mutovinia stella Minich 1992 ? Geryonichthys sp. Minikh 1998
Saurichthyiformes - Saurichthyidae	Saurichthys sp. Saurichthys sp. Agassiz 1834 ray-finned fish
Eurynotoidiformes - Eurynotoidiidae	? Isadia sp. ? Isadia sp. Minikh 1986 ray-finned fish
Ostracoda
-	Suchonellina bulloida, Suchonellina cf. anjugensis, Suchonellina ex gr. anjugensis, Suchonellina dispinosa, Suchonellina ex gr. dispinosa, Suchonellina ex gr. trapezoida, Suchonellina perelubica, Suchonellina pseudoinornata Suchonellina bulloida ostracod Suchonellina cf. anjugensis ostracod Suchonellina ex gr. anjugensis ostracod Suchonellina dispinosa ostracod Suchonellina ex gr. dispinosa ostracod Suchonellina ex gr. trapezoida ostracod Suchonellina perelubica ostracod = S. alija Suchonellina pseudoinornata ostracod = S. postparallela
Podocopida - Darwinulidae	Darwinula ? sima, Darwinula ? accuminata, Darwinula ? regia, Darwinula ? abscondita Darwinula ? sima ostracod Darwinula ? accuminata ostracod Darwinula ? regia ostracod Darwinula ? abscondita ostracod
-	Gerdalia ex gr. rixosa, Gerdalia ex gr. analoga, Gerdalia wetlugensis, ? Gerdalia sp. Gerdalia ex gr. rixosa ostracod Gerdalia ex gr. analoga ostracod Gerdalia wetlugensis ostracod ? Gerdalia sp. Belousova 1961 ostracod
Gastropoda
Neogastropoda - Marginellidae	? Marginella sp. ? Marginella sp. Lamarck 1799 margin shell
Bivalvia
Myalinida - Naiaditidae	Permianaia gusevi⁹ Permianaia gusevi⁹ Silantiev et al. 2018 clam

¹Battail and Surkov 2000; ²Bulanov 2003; ³Bulanov 2020; ⁴Bulanov and Yashina 2005; ⁵Gower and Sennikov 2000; ⁶Ivakhnenko 2008; ⁷Ivakhnenko 2011; ⁸Novikov et al. 2000; ⁹Silantiev et al. 2018