Twin Butte Creek (Cretaceous to of the United States)

Also known as Pteranodon harpyia holotype

Where: Logan County, Kansas (38.8° N, 100.9° W: paleocoordinates 44.1° N, 70.2° W)

• coordinate estimated from map

• local area-level geographic resolution

When: Smoky Hill Chalk Member (Niobrara Formation), Late/Upper Santonian to Late/Upper Santonian (85.8 - 70.6 Ma)

• Specimens probably collected between Marker Units 12 and 20 (Bennett 1991, 1994); upper part of member

• group of beds-level stratigraphic resolution

Environment/lithology: offshore; chalk

Size class: macrofossils

Collected by Cope in 1871; reposited in the AMNH

Primary reference: E. D. Cope. 1872. On two new ornithosaurians from Kansas. Proceedings of the American Philosophical Society 12:420-422 [M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 93225: authorized by Richard Butler, entered by Richard Butler on 08.01.2010, edited by Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

 Testudines - Toxochelyidae
Toxochelyidae indet.2 Baur 1895 sea turtle
Mandible and marginal scutes
 Testudines - Protostegidae
Protostega gigas3 Cope 1871 sea turtle
CM 1393, plastron, limbs, and skull
 Pterosauria -
Nyctosaurus sp.2 Marsh 1876 pterosaur
YPM 2287, limb-shaft fragments and two dorsal centra
Pteranodon longiceps1 Marsh 1876 pterosaur
CM 1539, CM "UNC A"
Ornithocheirus harpyia n. sp. Cope 1872 pterosaur
AMNH 1572, partial right metacarpal IV, proximal left wing phalanx 1, partial wing phalanx 2. At least two individuals