Cambridge Greensand [non-specific locality data] (Cretaceous of the United Kingdom)

Where: England, United Kingdom (52.2° N, 0.1° E: paleocoordinates 40.6° N, 2.8° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Cambridge Greensand Member (West Melbury Marly Chalk Formation), Late/Upper Albian (105.3 - 99.6 Ma)

• Remanie fauna - reworked from late Albian into early Cenomanian deposits. Faunal assemblage is late Albian in age (age used here). "Although the deposit itself is generally considered to be of late Cretaceous, Cenomanian age, its macrofossil assemblage is thought to be derived largely from the underlying early Cretaceous, Albian, Gault Formation, on the basis of derived ammonites associated with the vertebrate fossils. An early Cenomanian age for deposition of the Cambridge Greensand is based on an apparently autochthonous foraminiferan assemblage, but Morter and Wood (1983) could not rule out the possibility of a very late Albian age due to the presence of the bivalve Aucinella. The age of the derived vertebrates is now considered to be no older than the Callihoplites auritus ammonite subzone and most likely no younger than upper Stoliczkaia dispar Zone, and they are thus of late Albian age." (Martill & Unwin 2011, Cretaceous Research, 10.1016/j.cretres.2011.09.003).

• member-level stratigraphic resolution

Environment/lithology: open shallow subtidal; lag, glauconitic, micaceous, nodular, phosphatic, silty marl

Size class: macrofossils

Preservation: replaced with phosphate

Collection methods: salvage, mechanical,

• CAMSM: Sedgwick Museum, Cambridge

Primary reference: R. Owen. 1859. Supplement (No. I) to the Monograph on the Fossil Reptilia of the Cretaceous Formations. Palaeontographical Society Monographs 1-19 [R. Butler/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 94210: authorized by Richard Butler, entered by Richard Butler on 11.02.2010, edited by Bethany Allen, Roger Benson, Jonathan Tennant, Valentin Fischer, Patricia Holroyd and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Testudines - Cheloniidae
Lytoloma cantabrigiense n. sp.4 Lydekker 1889 sea turtle
BMNH 35178 (holotype), mandible
 Testudines - Pantestudinidae
Chelone jessoni n. sp.4 Lydekker 1889 turtle
BMNH 35186 (holotype), mandible
 Testudinata - Helochelydridae
Helochelys danubina3 von Meyer 1854 turtle
CAMSM B56424
 Testudinata -
Trachydermochelys phlyctaenus n. gen. n. sp.8 Seeley 1869 turtle
CAMSM 56340–6, seven peripheral plates
 Pterosauria -
Ornithocheirus enchorhynchus n. sp.9, Ornithocheirus xyphorhynchus n. sp.9, Ornithocheirus tenuirostris n. sp.9, Ornithocheirus carteri n. sp.9, Ornithocheirus eurygnathus n. sp.9, Ornithocheirus oxyrhinus n. sp.9, Ornithocheirus dentatus n. sp.9, Ornithostoma sedgwicki n. gen. n. sp.10, Ornithocheirus sp.11, "Pterodactylus simus n. sp." = Ornithocheirus simus5, "Ornithocheirus platyrhinus n. sp." = Ornithocheirus simus9, "Ornithocheirus platystomus n. sp." = Lonchodectes platystomus9, "Ornithocheirus microdon n. sp." = Lonchodectes microdon9, "Ornithocheirus oweni n. sp." = Lonchodectes microdon9, "Ornithocheirus huxleyi n. sp." = Lonchodectes microdon9, "Pterodactylus cuvieri" = Anhanguera cuvieri9, "Ornithocheirus colorhinus n. sp." = Anhanguera cuvieri9, "Ornithocheirus denticulatus n. sp." = Anhanguera cuvieri9, Ornithocheirus brachyrhinus n. sp.9, "Pterodactylus fittoni n. sp." = Camposipterus nasutus, "Ornithocheirus polyodon n. sp." = Camposipterus nasutus9, "Ornithocheirus nasutus n. sp." = Camposipterus nasutus7, Ornithocheirus scaphorhynchus n. sp.9, Ornithocheirus crassidens n. sp.9
Ornithocheirus enchorhynchus n. sp.9 Seeley 1870 pterosaur
CAMSM B 54444, anterior portion of the rostrum
Ornithocheirus xyphorhynchus n. sp.9 Seeley 1870 pterosaur
anterior portion of the rostrum (collection data could not be recovered)
Ornithocheirus tenuirostris n. sp.9 Seeley 1870 pterosaur
CAMSM B 54584, anterior portion of the rostrum
Ornithocheirus carteri n. sp.9 Seeley 1870 pterosaur
CAMSM B 54437, anterior portion of the rostrum
Ornithocheirus eurygnathus n. sp.9 Seeley 1870 pterosaur
CAMSM B54644, anterior fragment of an ?upper jaw
Ornithocheirus oxyrhinus n. sp.9 Seeley 1870 pterosaur
CAMSM B 54612, anterior fragment of an upper jaw
Ornithocheirus dentatus n. sp.9 Seeley 1870 pterosaur
CAMSM B 54544, anterior portion of the rostrum
Ornithostoma sedgwicki n. gen. n. sp.10 Seeley 1871 pterosaur
Ornithocheirus sp.11 Seeley 1869 pterosaur
CAMSM B54.890, possible new species
"Pterodactylus simus n. sp." = Ornithocheirus simus5 Owen 1861 pterosaur
CAMSM B54.428 (holotype), rostrum. Additional rostra and mandibular symphyses; CAMSM B54552, CAMSM B54429, CAMSM B54677, MANCH L.10832, and NHMUK PV 35412
"Ornithocheirus platyrhinus n. sp." = Ornithocheirus simus9 Owen 1861 pterosaur
"Ornithocheirus platystomus n. sp." = Lonchodectes platystomus9 Seeley 1870 pterosaur
"Ornithocheirus microdon n. sp." = Lonchodectes microdon9 Seeley 1869 pterosaur
CAMSM B54486, anterior portion of the rostrum
"Ornithocheirus oweni n. sp." = Lonchodectes microdon9 Seeley 1869 pterosaur
"Ornithocheirus huxleyi n. sp." = Lonchodectes microdon9 Seeley 1869 pterosaur
"Pterodactylus cuvieri" = Anhanguera cuvieri9 Bowerbank 1851 pterosaur
"Ornithocheirus colorhinus n. sp." = Anhanguera cuvieri9 Bowerbank 1851 pterosaur
"Ornithocheirus denticulatus n. sp." = Anhanguera cuvieri9 Bowerbank 1851 pterosaur
Anterior portion of the rostrum (?CAMSM B 54794)
Ornithocheirus brachyrhinus n. sp.9 Seeley 1870 pterosaur
CAMSM B54443, anterior portion of the rostrum
"Pterodactylus fittoni n. sp." = Camposipterus nasutus Seeley 1869 pterosaur
CAMSM B54.423 (holotype), anterior rostrum
"Ornithocheirus polyodon n. sp." = Camposipterus nasutus9 Seeley 1869 pterosaur
CAMSM B54440, anterior portion of the rostrum
"Ornithocheirus nasutus n. sp." = Camposipterus nasutus7 Seeley 1869 pterosaur
CAMSM B 54556 (holotype), anterior portion of the rostrum
Ornithocheirus scaphorhynchus n. sp.9 Seeley 1870 pterosaur
CAMSM B 54441, anterior portion of the rostrum
Ornithocheirus crassidens n. sp.9 Seeley 1870 pterosaur
 Pterosauria - Ornithocheiridae
"Ornithocheirus capito n. sp." = Coloborhynchus capito9, "Ornithocheirus reedi n. sp." = Coloborhynchus capito9, "Pterodactylus sedgwickii n. sp." = Coloborhynchus sedgwickii
"Ornithocheirus capito n. sp." = Coloborhynchus capito9 Seeley 1870 pterosaur
CAMSM B54.625 (holotype), rostrum.
"Ornithocheirus reedi n. sp." = Coloborhynchus capito9 Seeley 1870 pterosaur
"Pterodactylus sedgwickii n. sp." = Coloborhynchus sedgwickii Owen 1859 pterosaur
CAMSM B54.422 (holotype), anterior rostrum
 Pterosauria -
Pterodactylus woodwardi n. sp.5 Owen 1861 pterosaur
CAMSM B 54433a, anterior portion of the rostrum
 Sauropterygia -
Plesiosauria indet.8 de Blainville 1835 plesiosaur
Polyptychodon sp.8 Owen 1841 plesiosaur
Partial axial columns of two individuals, plus 103 teeth (p. 45)
Plesiosaurus pachyomus n. sp.8 Owen 1840 plesiosaur
Humerus, reputedly the type (p. 42). There is also a femur marked "? femur of the Pl. pachyomus type" (p. 44)
Plesiosaurus planus n. sp.6 Owen 1864 plesiosaur
p. 41, "series 5" [possibly more than one individual], "series 6", "series 7", mostly vertebrae
 Ichthyosauria -
Ichthyosauria indet.1 de Blainville 1835 ichthyosaur
Teeth NOT counted as valid specimens
 Ichthyosauria - Ophthalmosauridae
Ophthalmosauridae indet.1 Appleby 1956 ichthyosaur
Teeth NOT counted as valid specimens
Platypterygiinae indet.1 Arkhangelsky 2001 ichthyosaur
holotype of Ichthyosaurus bonneyi (CAMSM B58062, a femur); holotype of I. platymerus (CAMSM B58057, a humerus)
Platypterygius sp.2 von Huene 1922 ichthyosaur
CAMSM TN283* (rostrum and associated 112 teeth); CAMSM B42404_20* (basioccipital, centra); CAMSM B57939 (basioccipital); CAMSM B57940 (basioccipital); CAMSM B57944 (basioccipital); CAMSM B57959_60* (basioccipital, atlasaxis); CAMSM B58250_56* (quadrate, basioccipital, basisphenoid, supraoccipital, femur); CAMSM B75735 (basioccipital); CAMSM X50161 (basioccipital); CAMSM X50168 (basioccipital); CAMSM X50169 (basioccipital); CAMSM TN1729 partim (basioccipital); CAMSM TN1754 partim (basioccipital); CAMSM TN1755 partim (2 basioccipitals); NHMUK 35323 (basioccipital); several dozens of teeth, including CAMSM B57996_58009, CAMSM B58010 to B58027, CAMSM B58305_13; CAMSM B58379_87, CAMSM B76728_45, CAMSM TN1716 partim, CAMSM TN1778 partim; CAMSM TN1779 partim, NHMUK R625, NHMUK R133b partim, NHMUK R2336 partim (2 teeth), NHMUK 28110 partim, NHMUK 30253 partim, NHMUK 30254 (4 teeth), NHMUK 32406 partim, NHMUK 33242, NHMUK 35254 partim, NHMUK 35432_5, NHMUK 40358, NHMUK 41896, NHMUK 46381, NHMUK 47265_66* (teeth, centra), NHMUK 47269, IRSNB GS21, IRSNB GS25 to GS28, IRSNB GS32 to GS50, IRSNB GS53, IRSNB GS62; CAMSM B97401 partim (humerus morphotype 1); CAMSM B57987 (humerus morphotype 1); CAMSM B58043 (humerus morphotype 4); CAMSM B58048 (humerus morphotype 4); CAMSM B58056 (humerus morphotype 1); CAMSM B58057 (humerus morphotype 1; holotype of Ichthyosaurus platymerus); CAMSM B97401 partim (humerus morphotype 1); CAMSM TN1734 partim (humerus morphotype 4); CAMSM TN1751 partim (humerus morphotype 1); CAMSM TN1753 partim (one (humerus morphotype 1and one (humerus morphotype 4); CAMSM TN1757 partim (humerus morphotype 4); NHMUK R2342 partim (two humerus morphotype 4); CAMSM B58058 (femur); CAMSM B58060 (femur); CAMSM B58062 (femur; holotype of Ichthyosaurus bonneyi); CAMSM B58063_4 (femur); CAMSM B58361 (femur); CAMSM TN1749 partim (femur); CAMSM TN1748 partim (femur); CAMSM TN1757 partim (2 femora); NHMUK R23412 partim (femur); NHMUK R3510 (femur).
Sisteronia seeleyi n. gen. n. sp.2 Fischer et al. 2014 ichthyosaur
Holotype. CAMSM B58257_67, an incomplete specimen, including partial basicranium, scapula, humerus, and 5 centra; Referred material: CAMSM B57943 (basioccipital); CAMSM B57945 (basioccipital); CAMSM B57948 (basioccipital); CAMSM B57950 (basioccipital); CAMSM B57947 (basioccipital); CAMSM B57941 (basioccipital); CAMSM B57951 (basioccipital); CAMSM B57946 (basioccipital); CAMSM B57956 (basioccipital); CAMSM B57954 (basioccipital); CAMSM B58314 (basioccipital); CAMSM TN1727 partim (basioccipital); CAMSM TN1735 partim (6 basioccipitals); CAMSM TN1739 partim (basioccipital); CAMSM TN1751 partim (6 basioccipitals); CAMSM TN1753 partim (basioccipital); IRSNB GS54 (basioccipital); IRSNB GS61 (basioccipital); LEICT G107.1991 (basioccipital); NHMUK 44159 (basioccipital); NHMUK 44159a (basioccipital); CAMSM B57908 (opisthotic); CAMSM B58077_78 (2 opisthotics); CAMSM TN1753 partim (opisthotic); NHMUK R2348 (opisthotic); IRSNB GS10 (opisthotic); CAMSM B58091 (tooth); CAMSM B58092 (tooth); CAMSM TN1716 partim (numerous teeth); CAMSM TN1778 partim (numerous teeth); CAMSM TN1779 partim (numerous teeth); CAMSM B58390 (tooth); NHMUK R1923 (tooth); IRSNB GS23 (tooth); IRSNB GS24 (tooth); IRSNB GS55 to GS58 (teeth); CAMSM TN1755 partim (humerus); CAMSM TN1757 partim (humerus)
Ophthalmosaurinae indet.1 Baur 1887 ichthyosaur
NHMUK 32406 partim (tooth); NHMUK R16 partim (tooth); NHMUK 47268 (5 teeth); CAMSM B58042 (humerus); CAMSM B58045 (humerus); CAMSM B58050 (humerus); CAMSM B58053 (humerus); CAMSM B58055 (humerus); CAMSM TN1727 partim (humerus); CAMSM TN1755 partim (2 humeri); IRSNB GS3 (humerus); LEICT G65.1991 (humerus); NHMUK R2343 (3 humeri); NHMUK R4513 (2 humeri); NHMUK 35348 (humerus); NHMUK 43989 (humerus, holotype of Brachypterygius cantabrigiensis); IRSNB GS60 (anterior accessory epipodial element).
 Ichthyosauria -
Cetarthrosaurus walkeri n. gen. n. sp.2
Cetarthrosaurus walkeri n. gen. n. sp.2 Seeley 1869 ichthyosaur
Two propodials: CAMSM B58069, CAMSM X50170