Soyana (Iva Gora, PIN collection 94) (Permian of Russian Federation)

Where: Russian Federation (65.6° N, 43.6° E: paleocoordinates 33.2° N, 33.0° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size class: macrofossils

Collected by M. Edemskij in 1934

Primary reference: F. R. Schram. 1980. Miscellaneous late Paleozoic Malacostraca of the Soviet Union. Journal of Paleontology 54(3):542-547 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 96860: authorized by Matthew Clapham, entered by Matthew Clapham on 05.07.2010, edited by Jered Karr and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Malacostraca
 Isopoda - Palaeophreatoicidae
Palaeophreatoicus sojanensis n. gen. n. sp.
Palaeophreatoicus sojanensis n. gen. n. sp. Birshtein 1962 isopod
Insecta
 Eoblattida - Megakhosaridae
"Megakhosarina minuscula" = Blattokhosara minuscula1
"Megakhosarina minuscula" = Blattokhosara minuscula1 Aristov 2008 insect
PIN 94/131, 338 (and/or 238)
 Eoblattida - Bardapteridae
Olgaephilus emendatus18
Olgaephilus emendatus18 Storozhenko 1990 insect
PIN 94/120(121), 133(134)
 Reculida - Ivapteridae
Ivaptera sharovi2
Ivaptera sharovi2 Aristov 2009 winged insect
PIN 94/50,132, 580, 660, 748, 834,1044,1048
 Dicondylia - Kennedyidae
Kennedya ivensis9
Kennedya ivensis9 Nel et al. 2012 winged insect
PIN 94/2
 Palaeodictyoptera - Eukulojidae
Eukuloja cubitalis17, Eukuloja atrosumma17
Eukuloja cubitalis17 Martynov 1932 winged insect
PIN 94/3, 7, 8, 10, 11, 13, 20, 710, 737, 1164
Eukuloja atrosumma17 Sinitshenkova 1981 winged insect
PIN 94/935, 1158
 Raphidioptera - Parasialidae
Parasialis dissedens n. sp.15
Parasialis dissedens n. sp.15 Ponomarenko 1977 snakefly
PIN 94/1217, 820
 Dicondylia - Permithonidae
Permithonopsis obscura n. sp.12, "Permithonopsis proxima" = Permithonopsis ivensis12, "Permithonopsis triramosa n. sp." = Permegalomus maculipennis12
Permithonopsis obscura n. sp.12 Martynova 1952 winged insect
PIN 94/173(726)
"Permithonopsis proxima" = Permithonopsis ivensis12 Martynov 1932 winged insect
PIN 94/152
"Permithonopsis triramosa n. sp." = Permegalomus maculipennis12 Martynov 1932 winged insect
PIN 94/172
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis11
Kamopanorpa rotundipennis11 Martynov 1932 winged insect
PIN 94/771
 Mecoptera - Permochoristidae
"Permochorista maculipennis" = Petromantis maculipennis11, "Permochorista venosa" = Petromantis maculipennis11, Petromantis rossica11, "Petrochorista dubia" = Petromantis dubia11, Petrochorista lepida11, Petrochorista sojanensis n. sp.11, Agetopanorpa sp.14, "Agetochorista dubia" = Agetopanorpa dubia11, "Agetochorista tillyardiana" = Agetopanorpa tillyardiana11, "Agetochorista ornata" = Agetopanorpa ornata11, "Agetochorista splendida" = Agetopanorpa ornata11, "Agetochorista umbrata" = Agetopanorpa ornata11, "Agetochorista letopalae" = Agetopanorpa letopalae11
"Permochorista maculipennis" = Petromantis maculipennis11 Martynov 1932 scorpionfly
PIN 94/142, 167, 419, 592_589), 768, 1152, 1210
"Permochorista venosa" = Petromantis maculipennis11 Martynov 1932 scorpionfly
PIN 94/900, 1060, 1205
Petromantis rossica11 Handlirsch 1904 scorpionfly
PIN 94/439, 501, 531, 746, 758, 861, 911; 94/1230 ("Soyana")
"Petrochorista dubia" = Petromantis dubia11 Martynov 1932 scorpionfly
PIN 94/681, 730, 978, 1198
Petrochorista lepida11 Martynova 1948 scorpionfly
PIN 94/754
Petrochorista sojanensis n. sp.11 Martynova 1948 scorpionfly
PIN 94/1151
Agetopanorpa sp.14 Carpenter 1930 scorpionfly
PIN 94/175
"Agetochorista dubia" = Agetopanorpa dubia11 Martynov 1932 scorpionfly
PIN 94/179, 185, 540, 705 (2 wings), 827
"Agetochorista tillyardiana" = Agetopanorpa tillyardiana11 Martynova 1948 scorpionfly
PIN 94/893, 909
"Agetochorista ornata" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/149, 164, 168, 170, 174, 176, 516, 538, 778, 980, 1139, 1157, 1211, 1213, 1312
"Agetochorista splendida" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/165, 193, 595, 598, 671, 708, 709, 968, 1138; var. furcata 94/238
"Agetochorista umbrata" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/1204, 1213
"Agetochorista letopalae" = Agetopanorpa letopalae11 Martynova 1948 scorpionfly
PIN 94/1153
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata11, "Permopanorpa tenuis" = Permopanorpa martynovi11
"Permopanorpa angustata" = Martynopanorpa angustata11 Martynov 1932 scorpionfly
PIN 94/169, 515, 553, 620
"Permopanorpa tenuis" = Permopanorpa martynovi11 Willmann 1978 scorpionfly
PIN 94/544
 Dicondylia - Permembiidae
Letopaloptera carpenteriana n. sp.13
Letopaloptera carpenteriana n. sp.13 Martynova 1961 winged insect
PIN 94/561
 Dicondylia - Permosialidae
Permosialis latiformis12, "Permosialis cauleoides n. sp." = Permosialis paucinervis12, "Permosialis bifasciata" = Permosialis paucinervis12, "Permosialis marmorata n. sp." = Permosialis paucinervis12
Permosialis latiformis12 Martynova 1952 winged insect
PIN 94/536
"Permosialis cauleoides n. sp." = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/798
"Permosialis bifasciata" = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/497, 576, 1212
"Permosialis marmorata n. sp." = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/186(192), 721(676)
 Cnemidolestodea - Sylvabestiidae
Sojanopermula lucida n. gen. n. sp.19, "Permula edemskii n. sp." = Sojanopermula edemskii3
Sojanopermula lucida n. gen. n. sp.19 Storozhenko 1992 winged insect
PIN 94/117
"Permula edemskii n. sp." = Sojanopermula edemskii3 Aristov and Storozhenko 2011 winged insect
PIN 94/1027
 Paoliida - Paoliidae
Permotermopsis pectinata20, Stenaropodites sojanensis20, Sojanidelia nana n. sp.20, Sojanidelia striata20, Sojanidelia makarkini20, Sojanidelia fluctuosa20, Mezenalicula connata n. gen. n. sp.3
Permotermopsis pectinata20 Martynov 1937 winged insect
PIN 94/115(6)
Stenaropodites sojanensis20 Storozhenko 1992 winged insect
PIN 94/1126
Sojanidelia nana n. sp.20 Storozhenko 1992 winged insect
PIN 94/1122(1128), 1133
Sojanidelia striata20 Storozhenko 1992 winged insect
PIN 94/559(814), 780(1132)
Sojanidelia makarkini20 Storozhenko 1992 winged insect
PIN 94/370, 377
Sojanidelia fluctuosa20 Storozhenko 1992 winged insect
PIN 94/675(934)
Mezenalicula connata n. gen. n. sp.3 Aristov and Storozhenko 2011 winged insect
PIN 94/883
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis4
Camptoneurites soyanensis4 Aristov et al. 2010 ice crawler
PIN 94/29
 Glosselytrodea - Jurinidae
Jurina sojanensis10, Jurina reducta10, Jurina marginata10
Jurina sojanensis10 Martynov 1938 winged insect
PIN 94/555, 865; 864, 868 form radialis
Jurina reducta10 Martynov 1938 winged insect
PIN 94/549
Jurina marginata10 Martynov 1938 winged insect
PIN 94/352, 369, 403, 409, 432, 549, 807, 937, 961
 Glosselytrodea - Glosselytridae
Glosselytron multivenosum10
Glosselytron multivenosum10 Martynov 1938 winged insect
PIN 94/276
 Hypoperlida - Hypoperlidae
Boreopsocus danksae16
Boreopsocus danksae16 Shcherbakov 1994 winged insect
PIN 94/197, 1299, 662
 Hemiptera - Ingruidae
Ingruo lanceolata n. gen. n. sp.8
Ingruo lanceolata n. gen. n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/455
 Hemiptera - Scytinopteridae
Scytinoptera kokeni6, Scytinoptera reducta6
Scytinoptera kokeni6 Handlirsch 1904 true bug
PIN 94/481, 875
Scytinoptera reducta6 Martynov 1928 true bug
PIN 94/1179, 665, 469, 975, 472, 461, 408, 493, 922, 638, 245, 1251, 395, 881, 257, 585, 431, 784, 731, 912
 Hemiptera - Coleoscytidae
Coleoscyta occallata n. sp.8
Coleoscyta occallata n. sp.8 Becker-Migdisova 1960 planthopper
PIN 94/1178(482)
 Hemiptera - Prosbolidae
Sojanoneura bimaculata5, Sojanoneura furcatula5, Sojanoneura triangularis5, Sojanoneura kazanensis5, Prosbole permocicadoides5, Prosbole indistincta n. sp.5, Prosbole sojanensis5, Prosbole incerta n. sp.5, Prosbole brevialata n. sp.5, Prosbole longialata n. sp.5, Prosbole ivagorae5, Permocicada nigronervosa5, Permocicada simulatrix5, Permocicada integra n. sp.5, Permocicada triramosa n. sp.5, Permocicada sublata n. sp.5, Permocicada pusilla n. sp.5, Permocicada idelensis5, Permocicadopsis angustata5, Permocicadopsis aberrans5
Sojanoneura bimaculata5 Martynov 1935 true bug
PIN 94/458, 639ab, 921, 862
Sojanoneura furcatula5 Martynov 1935 true bug
PIN 94/624(586), 579, 810
Sojanoneura triangularis5 Martynov 1935 true bug
PIN 94/953, 1194, 640
Sojanoneura kazanensis5 Zalessky 1929 true bug
PIN 94/466(452), 933
Prosbole permocicadoides5 Becker-Migdisova 1940 true bug
PIN 94/943, 253
Prosbole indistincta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/942
Prosbole sojanensis5 Martynov 1935 true bug
PIN 94/1174(467)
Prosbole incerta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/902
Prosbole brevialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/763(1190)
Prosbole longialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1176
Prosbole ivagorae5 Becker-Migdisova 1940 true bug
PIN 94/255(475)
Permocicada nigronervosa5 Martynov 1928 true bug
PIN 94/249
Permocicada simulatrix5 Martynov 1935 true bug
PIN 94/465, 1189
Permocicada integra n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/906ab
Permocicada triramosa n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1196
Permocicada sublata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1055(1043), 179
Permocicada pusilla n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/688
Permocicada idelensis5 Zalessky 1929 true bug
PIN 94/250(451)
Permocicadopsis angustata5 Martynov 1935 true bug
PIN 94/645, 964; 468, 1047, 389, 886(var. typica)
Permocicadopsis aberrans5 Martynov 1935 true bug
PIN 94/462, 8885
 Hemiptera - Prosbolopseidae
? Scytoneura sp.8, Scytoneura elliptica8, Anaprosbole ivensis8, Prosbolopsis ovalis8
? Scytoneura sp.8 Martynov 1935 true bug
PIN 94/641
Scytoneura elliptica8 Martynov 1935 true bug
PIN 94/243, 450, 473, 480, 483, 577, 596, 1191, 1192, 1283
Anaprosbole ivensis8 Becker-Migdisova 1960 true bug
PIN 94/464
Prosbolopsis ovalis8 Martynov 1935 true bug
PIN 94/420, 254, 251, 1184, 171
 Hemiptera - Boreoscytidae
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7 Becker-Migdisova 1949 true bug
PIN 94/1017, 775
 Hemiptera - Archescytinidae
Sojanoscytina ramosa n. sp.8, Sojanoscytina ivagorae n. sp.8, Archescytina fusca8, "Sojanoscytina tetraclada" = Archescytina tetraclada8
Sojanoscytina ramosa n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/1182
Sojanoscytina ivagorae n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/887
Archescytina fusca8 Becker-Migdisova 1960 true bug
PIN 94/474, 107
"Sojanoscytina tetraclada" = Archescytina tetraclada8 Martynov 1933 true bug
PIN 94/1185
Lingulata
 Lingulida - Lingulidae
Lingula orientalis, Lingula credneri
Lingula credneri Geinitz 1848
Rhynchonellata
 Rhynchonellida - Rhynchoporidae
Rhynchopora geinitziana
Rhynchopora geinitziana de Verneuil 1845
Strophomenata
 Productida - Linoproductidae
"Productus cancrini" = Cancrinella cancrini
"Productus cancrini" = Cancrinella cancrini de Verneuil 1845
Bivalvia
 Ostreida - Pinnidae
Aviculopinna ivanisciana
 Pectinida - Streblochondriidae
"Pecten sericeum" = Streblochondria sericea
"Pecten sericeum" = Streblochondria sericea de Verneuil 1845 scallop
 Pectinida - Pseudomonotidae
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria von Schlotheim 1816 scallop
1Aristov 2008; 2Aristov 2009; 3Aristov and Storozhenko 2011; 4Aristov et al. 2010; 5Becker-Migdisova 1940; 6Becker-Migdisova 1948; 7Becker-Migdisova 1949; 8Becker-Migdisova 1960; 9Felker 2023; 10Martynov 1938; 11Martynova 1948; 12Martynova 1952; 13Martynova 1961; 14Novokshonov 1994; 15Ponomarenko 1977; 16Shcherbakov 1994; 17Sinitshenkova 1981; 18Storozhenko 1990; 19Storozhenko 1992; 20Storozhenko 1992