Soyana (Iva Gora, PIN collection 94) (Permian of Russian Federation)

Where: Russian Federation (65.6° N, 43.6° E: paleocoordinates 33.2° N, 33.0° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size class: macrofossils

Collected by M. Edemskij in 1934

Primary reference: F. R. Schram. 1980. Miscellaneous late Paleozoic Malacostraca of the Soviet Union. Journal of Paleontology 54(3):542-547 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 96860: authorized by Matthew Clapham, entered by Matthew Clapham on 05.07.2010, edited by Jered Karr and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hypoperlida - Hypoperlidae
Boreopsocus danksae15 Shcherbakov 1994 winged insect
PIN 94/197, 1299, 662
 Hemiptera - Coleoscytidae
Coleoscyta occallata n. sp.8 Becker-Migdisova 1960 planthopper
PIN 94/1178(482)
 Hemiptera - Scytinopteridae
Scytinoptera kokeni6 Handlirsch 1904 true bug
PIN 94/481, 875
Scytinoptera reducta6 Martynov 1928 true bug
PIN 94/1179, 665, 469, 975, 472, 461, 408, 493, 922, 638, 245, 1251, 395, 881, 257, 585, 431, 784, 731, 912
 Hemiptera - Ingruidae
Ingruo lanceolata n. gen. n. sp.8
Ingruo lanceolata n. gen. n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/455
 Hemiptera - Archescytinidae
Archescytina fusca8 Becker-Migdisova 1960 true bug
PIN 94/474, 107
"Sojanoscytina tetraclada" = Archescytina tetraclada8 Martynov 1933 true bug
PIN 94/1185
Sojanoscytina ramosa n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/1182
Sojanoscytina ivagorae n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/887
 Hemiptera - Boreoscytidae
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7 Becker-Migdisova 1949 true bug
PIN 94/1017, 775
 Hemiptera - Prosbolopseidae
Anaprosbole ivensis8 Becker-Migdisova 1960 true bug
PIN 94/464
? Scytoneura sp.8 Martynov 1935 true bug
PIN 94/641
Scytoneura elliptica8 Martynov 1935 true bug
PIN 94/243, 450, 473, 480, 483, 577, 596, 1191, 1192, 1283
Prosbolopsis ovalis8 Martynov 1935 true bug
PIN 94/420, 254, 251, 1184, 171
 Hemiptera - Prosbolidae
Permocicada simulatrix5 Martynov 1935 true bug
PIN 94/465, 1189
Permocicada nigronervosa5 Martynov 1928 true bug
PIN 94/249
Permocicada sublata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1055(1043), 179
Permocicada triramosa n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1196
Permocicada integra n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/906ab
Permocicada idelensis5 Zalessky 1929 true bug
PIN 94/250(451)
Permocicada pusilla n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/688
Permocicadopsis angustata5 Martynov 1935 true bug
PIN 94/645, 964; 468, 1047, 389, 886(var. typica)
Permocicadopsis aberrans5 Martynov 1935 true bug
PIN 94/462, 8885
Sojanoneura bimaculata5 Martynov 1935 true bug
PIN 94/458, 639ab, 921, 862
Sojanoneura furcatula5 Martynov 1935 true bug
PIN 94/624(586), 579, 810
Sojanoneura triangularis5 Martynov 1935 true bug
PIN 94/953, 1194, 640
Sojanoneura kazanensis5 Zalessky 1929 true bug
PIN 94/466(452), 933
Prosbole incerta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/902
Prosbole indistincta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/942
Prosbole sojanensis5 Martynov 1935 true bug
PIN 94/1174(467)
Prosbole longialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1176
Prosbole ivagorae5 Becker-Migdisova 1940 true bug
PIN 94/255(475)
Prosbole brevialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/763(1190)
Prosbole permocicadoides5 Becker-Migdisova 1940 true bug
PIN 94/943, 253
 Dicondylia - Permosialidae
Permosialis latiformis11 Martynova 1952 winged insect
PIN 94/536
Permosialis bifasciata11 Martynov 1932 winged insect
PIN 94/497, 576, 1212
Permosialis cauleoides n. sp.11 Martynova 1952 winged insect
PIN 94/798
Permosialis marmorata n. sp.11 Martynova 1952 winged insect
PIN 94/186(192), 721(676)
 Cnemidolestodea - Sylvabestiidae
Sojanopermula lucida n. gen. n. sp.18, "Permula edemskii n. sp." = Sojanopermula edemskii3
Sojanopermula lucida n. gen. n. sp.18 Storozhenko 1992 winged insect
PIN 94/117
"Permula edemskii n. sp." = Sojanopermula edemskii3 Aristov and Storozhenko 2011 winged insect
PIN 94/1027
 Paoliida - Paoliidae
Stenaropodites sojanensis19 Storozhenko 1992 winged insect
PIN 94/1126
Sojanidelia fluctuosa19 Storozhenko 1992 winged insect
PIN 94/675(934)
Sojanidelia makarkini19 Storozhenko 1992 winged insect
PIN 94/370, 377
Sojanidelia striata19 Storozhenko 1992 winged insect
PIN 94/559(814), 780(1132)
Sojanidelia nana n. sp.19 Storozhenko 1992 winged insect
PIN 94/1122(1128), 1133
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis4 Aristov et al. 2010 ice crawler
PIN 94/29
 Dicondylia - Permembiidae
Letopaloptera carpenteriana n. sp.12 Martynova 1961 winged insect
PIN 94/561
 Glosselytrodea - Glosselytridae
Glosselytron multivenosum9 Martynov 1938 winged insect
PIN 94/276
 Glosselytrodea - Jurinidae
Jurina reducta9 Martynov 1938 winged insect
PIN 94/549
Jurina sojanensis9 Martynov 1938 winged insect
PIN 94/555, 865; 864, 868 form radialis
Jurina marginata9 Martynov 1938 winged insect
PIN 94/352, 369, 403, 409, 432, 549, 807, 937, 961
 Mecoptera - Permochoristidae
Petromantis rossica10, "Permochorista maculipennis" = Petromantis maculipennis10, "Permochorista venosa" = Petromantis maculipennis10, "Petrochorista dubia" = Petromantis dubia10, Petrochorista sojanensis n. sp.10, Petrochorista lepida10, Agetopanorpa sp.13, "Agetochorista letopalae" = Agetopanorpa letopalae10, "Agetochorista ornata" = Agetopanorpa ornata10, "Agetochorista umbrata" = Agetopanorpa ornata10, "Agetochorista splendida" = Agetopanorpa ornata10, "Agetochorista dubia" = Agetopanorpa dubia10, "Agetochorista tillyardiana" = Agetopanorpa tillyardiana10
Petromantis rossica10 Handlirsch 1904 scorpionfly
PIN 94/439, 501, 531, 746, 758, 861, 911; 94/1230 ("Soyana")
"Permochorista maculipennis" = Petromantis maculipennis10 Martynov 1932 scorpionfly
PIN 94/142, 167, 419, 592_589), 768, 1152, 1210
"Permochorista venosa" = Petromantis maculipennis10 Martynov 1932 scorpionfly
PIN 94/900, 1060, 1205
"Petrochorista dubia" = Petromantis dubia10 Martynov 1932 scorpionfly
PIN 94/681, 730, 978, 1198
Petrochorista sojanensis n. sp.10 Martynova 1948 scorpionfly
PIN 94/1151
Petrochorista lepida10 Martynova 1948 scorpionfly
PIN 94/754
Agetopanorpa sp.13 Carpenter 1930 scorpionfly
PIN 94/175
"Agetochorista letopalae" = Agetopanorpa letopalae10 Martynova 1948 scorpionfly
PIN 94/1153
"Agetochorista ornata" = Agetopanorpa ornata10 Martynov 1932 scorpionfly
PIN 94/149, 164, 168, 170, 174, 176, 516, 538, 778, 980, 1139, 1157, 1211, 1213, 1312
"Agetochorista umbrata" = Agetopanorpa ornata10 Martynov 1932 scorpionfly
PIN 94/1204, 1213
"Agetochorista splendida" = Agetopanorpa ornata10 Martynov 1932 scorpionfly
PIN 94/165, 193, 595, 598, 671, 708, 709, 968, 1138; var. furcata 94/238
"Agetochorista dubia" = Agetopanorpa dubia10 Martynov 1932 scorpionfly
PIN 94/179, 185, 540, 705 (2 wings), 827
"Agetochorista tillyardiana" = Agetopanorpa tillyardiana10 Martynova 1948 scorpionfly
PIN 94/893, 909
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata10, "Permopanorpa tenuis" = Permopanorpa martynovi10
"Permopanorpa angustata" = Martynopanorpa angustata10 Martynov 1932 scorpionfly
PIN 94/169, 515, 553, 620
"Permopanorpa tenuis" = Permopanorpa martynovi10 Willmann 1978 scorpionfly
PIN 94/544
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis10 Martynov 1932 winged insect
PIN 94/771
 Raphidioptera - Parasialidae
Parasialis dissedens n. sp.14 Ponomarenko 1977 snakefly
PIN 94/1217, 820
 Dicondylia - Permithonidae
"Permithonopsis proxima" = Permithonopsis ivensis11, Permithonopsis obscura n. sp.11, "Permithonopsis triramosa n. sp." = Permegalomus maculipennis11
"Permithonopsis proxima" = Permithonopsis ivensis11 Martynov 1932 winged insect
PIN 94/152
Permithonopsis obscura n. sp.11 Martynova 1952 winged insect
PIN 94/173(726)
"Permithonopsis triramosa n. sp." = Permegalomus maculipennis11 Martynov 1932 winged insect
PIN 94/172
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma16 Sinitshenkova 1981 winged insect
PIN 94/935, 1158
Eukuloja cubitalis16 Martynov 1932 winged insect
PIN 94/3, 7, 8, 10, 11, 13, 20, 710, 737, 1164
 Reculida - Ivapteridae
Ivaptera sharovi2 Aristov 2009 winged insect
PIN 94/50,132, 580, 660, 748, 834,1044,1048
 Eoblattida - Permotermopsidae
Mezenalicula connata n. gen. n. sp.3 Aristov and Storozhenko 2011 insect
PIN 94/883
Permotermopsis pectinata19 Martynov 1937 insect
PIN 94/115(6)
 Eoblattida - Megakhosaridae
"Megakhosarina minuscula" = Blattokhosara minuscula1
"Megakhosarina minuscula" = Blattokhosara minuscula1 Aristov 2008 insect
PIN 94/131, 338 (and/or 238)
 Eoblattida - Bardapteridae
Olgaephilus emendatus17 Storozhenko 1990 insect
PIN 94/120(121), 133(134)
Malacostraca
 Isopoda - Palaeophreatoicidae
Palaeophreatoicus sojanensis n. gen. n. sp. Birshtein 1962 isopod
Bivalvia
 Pectinida - Streblochondriidae
"Pecten sericeum" = Streblochondria sericea
"Pecten sericeum" = Streblochondria sericea de Verneuil 1845 scallop
 Pectinida - Pseudomonotidae
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria von Schlotheim 1816 scallop
 Ostreida - Pinnidae
Strophomenata
 Productida - Linoproductidae
"Productus cancrini" = Cancrinella cancrini
"Productus cancrini" = Cancrinella cancrini de Verneuil 1845
Rhynchonellata
 Rhynchonellida - Rhynchoporidae
Rhynchopora geinitziana de Verneuil 1845
Lingulata
 Lingulida - Lingulidae