Insecta - Coleoptera - Carabidae
Alternative spellings: Harpalidae, Harpalii
Synonym: Licininae Bonelli 1810
Full reference: F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques) 1-58
Parent taxon: Carabidae according to C. Y. Cai et al. 2022
See also Bouchard et al. 2011, Kirichenko-Babko et al. 2021, Lawrence and Newton 1995, LeConte 1861, Liu et al. 2015, Tamutis et al. 2011 and Webster 2016
Sister taxa: Antemnacrassa, Apotomopterus, Ataktosites, Bembidium, Brachininae, Brachinites, Broscinae, Carabidium, Carabinae, Carabites, Carabites, Carabopteron, Cavicarabus, Celeripedes, Cicindelites, Conjunctiinae, Cratacanthus florissantensis, Elaphrinae, Elaphrotites, Fangshania, Feronites, Fonsecacarabus, Fossores, Gehringiinae, Glenopterus, Hadrocephalus, Harpalomimes, Hebeicarabus, Lithocantharis, Loricera exita, Loricerinae, Meileyingia, Metagrilium, Migadopinae, Nebriinae, Nebrorabus, Neothanes, Notokistus, Omophroninae, Paussinae, Platyninae, Procarabites, Protopaussinae, Protorabinae, Rhopalochlaenius, Rhysodinae, Scaritinae, Siagoninae, Sinocarabus, Stereocerus, Sunocarabus, Trechinae, Trechoides, Trechus fractus, Tremembecarabus, Wongyekokia, Wuchangicarabus, Yunnanocarabus
Subtaxa: Abacetini Aethocarabus Chlaeniini Dryptini Galeritini Harpalini Helluonini Lebiini Licinini Odacanthini Oodini Panagaeini Pseudomorphini Pterostichini Sinis Sphodrini Zabrini Zuphiini
Type: Harpalus
Ecology: