Distribution:
• Quaternary of Bolivia (1 collection), Brazil (4), Canada (2: Northwest Territories, Quebec), Chile (1), Greece (1), Greenland (3), Italy (1), Japan (1), the Russian Federation (1), Spain (1), Turkmenistan (1), United States (5: California, Nevada, Pennsylvania, Virginia), Venezuela (1)
• Blancan of United States (1: California)
• Pliocene of Greece (1), Kazakhstan (1), Romania (1), South Africa (1)
• Hemphillian of United States (1: Washington)
• Miocene to Pliocene of Turkey (1)
• Miocene of Canada (2: Northwest Territories, Yukon), China (1), France (1), Germany (4), Greece (1), Italy (2), Kazakhstan (1), Kenya (1), Pakistan (12), the Russian Federation (1), Slovakia (1), Switzerland (2), United States (14: California, Florida, Louisiana, Montana, North Carolina, Oklahoma, South Dakota, Texas, Utah)
• Arikareean of United States (1: California)
• Oligocene to Miocene of Kazakhstan (1), Pakistan (1)
• Oligocene of China (1), Egypt (1), Pakistan (2), Serbia and Montenegro (3), Thailand (1), United States (1: South Dakota)
• Eocene to Oligocene of China (1)
• Eocene of China (2), France (7), Kazakhstan (1), Kyrgyzstan (1), Pakistan (2), Serbia and Montenegro (1), Spain (5), Turkey (1), the United Kingdom (2), United States (22: California, Utah, Wyoming)
• Cenozoic of Mongolia (1)
Total: 131 collections including 132 occurrences