Distribution:
• Quaternary of Italy (1 collection)
• Miocene of Austria (2), Germany (1), Greece (2), Poland (1)
• Eocene of the United Kingdom (1)
• Cretaceous of Italy (17), Oman (1)
• Jurassic of Albania (3), United States (3: Alaska, Oregon)
Total: 32 collections including 38 occurrences
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