Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240
See also Ponomarenko 1969
Sister taxa: none
Type specimen: PIN 1700/3204, an exoskeleton. Its type locality is Chekarda, Sylva River (PIN collection 1700), which is in a Kungurian deltaic siliciclastic in the Koshelevka Formation of the Russian Federation.
Distribution: found only at Chekarda, Sylva River (PIN collection 1700)
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.